8b. var. pungens
Stigmas mostly 2. Fruits flattened and
unequally biconvex in cross-section (flat on 1 side, rounded on the other).
May–September.
Scattered nearly throughout Missouri, but more common south of the Missouri River (U.S. and Canada south to South
America and the Caribbean Islands; Europe). Margins of ponds, lakes, sloughs,
and ditches, gravel bars, and stream banks, fens, and marshes.
Other characters included by Koyama (1963)
and Beetle (1947) do not correlate with the separation of bicarpellate from
tricarpellate plants. Of the two varieties, var. pungens is by far the
most common in Missouri. In the United States var. pungens is most
common in the eastern half, particularly east of the Appalachian Mountains,
whereas var. longispicatus predominates in the western half. Missouri is part of a broad zone of geographic overlap between the two varieties, and
intermediates (with nearly equal numbers of bicarpellate and tricarpellate
florets) have been recorded from areas where both taxa occur. The character of
fruit shape can be relatively subtle and easily misinterpreted.