66. Carex molestiformis Reznicek
& P. Rothr.
Map 182
Plants
with short, inconspicuous rhizomes, forming tufts or clumps. Vegetative
stems shorter than the flowering stems and developing late in the season, the
leaves clustered near the tip. Flowering stems 25–120 cm long, much longer than
the leaves. Leaves with well-developed blades mostly 3–5 per
flowering stem. Leaf blades 2–30 cm long, 1.5–4.0 mm wide, green. Leaf
sheaths often extended past the insertion point of the leaf blade, the ventral
side white to light yellow, papery, the tip truncate to somewhat concave, the ligule mostly wider than long
and U-shaped. Inflorescence straight or somewhat bent immediately above the lowermost
spike, the 2–5 spikes densely overlapping along the axis, the lowermost spike
sometimes more loosely spaced. Spikes 7–17 mm long, 5–10 mm wide, the pistillate portion circular to broadly ovate or obovate in outline, rounded at both ends, with usually
numerous perigynia with appressed
to ascending or somewhat spreading tips, the staminate portion usually
inconspicuous. Scales 2.5–5.0 mm long, shorter and narrower than, but not
hidden by the perigynia, narrowly ovate, rounded to
sharply pointed, white to light yellowish brown with a green or pale brown
midrib. Perigynia 4.0–5.4 mm long, 2.4–3.2 mm wide,
1.3–1.9 times as long as wide, flat to slightly concave on the ventral side and
somewhat convex on the dorsal side, the main body as long as wide or slightly
longer than wide, broadly ovate-elliptic, widest at or below the middle,
broadly winged to the base, rounded gradually to a beak with toothed or
roughened margins, the wing extending to the tip of the beak, the ventral and
dorsal surfaces lacking papillae, sharply 4–6-nerved on the ventral surface and
finely 6–11-nerved on the dorsal surface, greenish brown to light brown. Fruits
1.6–2.0 mm long, 1.4–1.6 mm wide, 1.0–1.3 times as long as wide, broadly oblong
to nearly circular in outline, light brown to brown. 2n=74. May–July.
Uncommon
in the eastern half of the Ozark Division (endemic to the Ozarks of Arkansas, Missouri, and Oklahoma).
Banks of streams, rivers, and spring branches, bottomland
forests, open bottoms of ravines in mesic upland forests,
fens, and shaded dolomite bluffs.
This recently described
Ozark endemic (Reznicek and Rothrock,
1997) refers to a series of populations that consistently have perigynia at the large end of the range of variation for C.
molesta, and which differ from that species in
the shape of their fruits. The plants also tend to have stems that are stouter
toward the tip. Some specimens of C. molestiformis
originally were determined as C. brevior,
because the spikes with more ascending perigynia
superficially resemble that species. However, C. brevior
has the perigynia nerveless or nearly so.