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  • Reference article Robbrecht, E. & J. F. Manen. 2006. The major evolutionary lineages of the coffee family (Rubiaceae, angiosperms). Combined analysis (nDNA and cpDNA) to inferthe position of Coptosapelta and Luculia, and supertree construction based on rbcL, rps16, trnL-trnF and atpB-rbcL data. A new classification in two subfamilies, Cinchonoideae and Rubioideae. Syst. & Geogr. Pl. 76: 85–146.
  • Reference article Taylor, C. M. 2010-2025. Studies in Tropical Rubiaceae.
 
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