Lateral nectar glands various; middle glands present or absent. Hairs simple or branched, often 2-3-partite appressed. Sepals erect, rarely spreading, inner often saccate or subsaccate at base. Stamen filaments not appendaged, free, rarely those of the inner four connate in pairs; anthers dithecous (rarely monothecous), oblong, obtuse. Ovary sessile or shortly stipitate; stigma ± bilobed. Fruits usually long (siliquae), rarely short, dehiscent or indehiscent, sometimes breaking transversely into 1-seeded parts. Seeds not winged, rarely obscurely or slightly winged; radicle incumbent or rarely accumbent.