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(Last Modified On 1/28/2013)
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(Last Modified On 1/28/2013)
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Subspecific
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ARENARIA LANUGINOSA subSP. guatemalensis (Standl. & Steyerm.) J. Duke
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Synonym
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Arenaria guatemalensis Standl. & Steyerm. in Field Mus. Bot. 23:50. 1944.
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Description
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Herbaceous puberulent laxly spreading perennials. Leaves opposite, lanceolate, apically attenuate and mucronulate, basally subsessile and attenuate, 10-40 mm. long, 2-6 mm. broad, irregularly puberulent with a whitish indumentum, often ciliate on the margins. Flowers solitary in the axils, the puberulent ebracteate pedicels 10-40 mm. long, usually exceeding the subtending leaves. Sepals lanceolate to ovate, attenuate, strongly puberulent on the costa, 4-6 mm. long; petals 5, obovate, entire, 5-8 mm. long; stamens 10, the filaments 3-6 mm. long; ovary sessile or short-stipitate, globose or ovoid; styles 2-4. Capsule ovoid, 5-8 mm. long, the 3 valves deeply emarginate; seeds numerous, on basal placentae, cochleate, dark reddish brown to black, smooth or minutely tuberculate, ca. 0.75 mm. broad.
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Distribution
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Mexico to Panama, at moderate elevations.
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Specimen
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CHIRIQUf: rain forest, Bajo Chorro, Boquete District, 6000 ft., Davidson 256; vicinity of Finca Lerida, 1750 m., Woodson d Schery 214; vicinity of Casita Alta, Volcain Chiri- qui, ca. 1500-2000 m., Woodson, Allen & Seibert 89; valley of the upper Rio Chiriqui Viejo, vicinity of Monte Lirio, 1300-1900 m., Seibert 297; in grassy sunny places, vicinity of Cerro Punta, 1500-2000 m., Seibert 260.
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Note
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Other variants of the lanuginosa complex are found in Mexico and Central America and could conceivably occur in Panama. In the subspecies guatemalensis, the variety ensifolia, with linear leaves and exerted petals, is so far reported only from southern Mexico. Characterized by ovate leaves, cuneate to cordate leaf bases and exerted petals, the variant currently passing as A. megalantha (Rohrb.) F. N. Williams would probably better be treated as a lanuginosa subspecies, ranging front southern Mexico to Guatemala. The original description of A. guaternalensis would have been more edifying had it been contrasted with A. lanuginosa instead of A. megalantha, for although intergrades occur between all three, they are more frequent and perplexing between the first two. The relative constancy of these three taxa, where their ranges are distinct, coupled with limited intergradations near the commissures of their ranges, seems to justify their relegation to the subspecific level, in accord with the heirarchy established by Maguire (in Am. Midl. Nat. 46:493. 1951) in his monograph of the North American species of Arenaria.
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Reference
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Am. Midl. Nat. 46:493. 1951
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