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(Last Modified On 7/31/2013)
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(Last Modified On 7/31/2013)
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Species
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Gurania megistantha Donnell Smith
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PlaceOfPublication
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Bot. Gaz. (Crawfordsville) 33: 251. 1902.
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Note
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TYPE: Costa Rica, Smith 6649 (US, holotype).
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Synonym
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Gurania suberosa Standley, J. Wash. Acad. Sci. 15: 461. 1925. TYPE: Panama, Standley 31461 (US, holotype).
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Description
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Dioecious, tendriled lianas; juvenile stems 5- to 10-sulcate, pilose, older stems woody, with thick, ridged, corky periderm. Leaves simple, broadly ovate to sub- orbicular, shallowly to deeply 3-lobate, rarely 5-lobate or entire, 10-25 cm long, 9-20 cm wide, the apex acuminate, the base cordate, the margin minutely and remotely denticulate, chartaceous, the upper surface minutely pustulate, the up- per and the lower surfaces pilose, especially on the veins; petioles 4-11 cm long, slender, striate, pilose, the tendrils simple, pilose or glabrate. Staminate flowers in sessile or subsessile, umbelloid racemes at the nodal regions on the old leafless stems, less commonly on short herbaceous branches arising from the old wood or the young stems, the rachis strigose, puberulent, or glabrate; peduncles 1-2 (-12) cm long; pedicels 1-3 cm long; calyx orange, tubular with a bulbous base, 1.5-3.5 cm long, the outer surface puberulent or glabrate, the inner surface densely papillose puberulent, lobed up to ca. ? its length, the lobes linear, spreading; corolla yellow, equalling or slightly exceeding the calyx, the lobes lanceolate, acute, densely papillose puberulent on both the surfaces; stamens 2, free, attached to the hypanthium wall, subsessile, the anthers linear ca. 2 cm long, replicate at the base, the appendix linear triangulate, papillose, 8-10 cm long. Pistillate flowers in sessile or subsessile, umbelloid racemes on leafless branches on the upper part of the stem; calyx and corolla as in the staminate flowers; ovary oblong, glabrate, ovules numerous, horizontal, the styles 2, con- nate, thick, the stigmas 2, linear lanceolate, the inner surface papillose puberu- lent and medially grooved, ca. 1/? the length of the style column. Fruits borne in tight clusters, pale green with light spots, elliptic 4-5 cm long, 1.5-2 cm wide, longitudinally grooved, old flowers persisting; seeds white, elliptic ovoid, com- pressed, 6-9 mm long, 3-4 mm wide, the testa rugulose.
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Habit
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lianas
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Note
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Gurania megistantha is a species of tropical wet forest at elevations up to 1,000 m. It is also known from Costa Rica. The bright orange staminate flow- ers, with a long tubular hypanthium and short calyx lobes, usually borne on woody, corky, naked branches, make this one of the easiest of the Panamanian Gurania species to identify.
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Elevation
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up to 1,000 m.
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Specimen
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BOCAS DEL TORO: Region of Almirante, Buena Vista, Cooper 215 (F). Quebrado Huron, Kirkbride & Duke 423 (MO). Fish Creek Mountains, Chiriqui Lagoon, Wedel 2202 (GH, MO), 2297 (MO), 2372 (GH, MO, US). CANAL ZONE: Mojinga Swamp near mouth of Rio Chagres, Allen 906 (GH, MO, NY). Telephone trail between Rio Indio Hydrographic Station and the natural bridge of Rio Puente, Dodge & Allen 17477 (MO). Rio Providencia, 3 km SE of Achiote near W border of Canal Zone, Gentry & Nee 8671 (MO, USF). Rio Indio de Gatun, Goldman 2778 (US). Madden Forest Preserve, Las Cruces Trail and Highway, Lewis et al. 5305 (MO). Quebrada San Antonio, 2.0-3.5 km N of Gamboa, Nee 9466 (MO, USF). BARRO COLORADO ISLAND: Aviles 28 (F); Bailey 205 (F); Brown 102a (F); CGoat 5782, 8021, 12217, 12933, 14105A, 14554 (all MO); Foster 654 (DUKE); Hunnewell 16463 (GH); Shat- tuck 566 (F, MO); Standley 40916 (US); Wetmore et al. 10 (GH), 10A (F, GH); Zetek 3845 (F). COCLE: El Valle de Anton, Allen 1807 (F, GH, MO, US). Cerro Pilon, Croat 22938 (F, MO, US). El Valle de Anton, Dwyer 10529 (MO). Cerro Pilon, Dwyer & Lalla- thin 8640 (MO). El Valle de Anton, Lewis et al. 1778 (MO). Boca del Toabre at confluence of Rio Toabre and Rio Cocle del Norte, Lewis et al. 5574 (MO). COLON: 1-3 mi W of Portobelo, Gentry 1760 (F, MO). Ca. 0.5 km fromn Rio Buenaventura near Portobelo, Ken- nedy & Gra 2246 (F, MO, US). 10 mi SW of Portobelo and 2-4 mi from coast, Liesner 1036 (MO). Rio Lagarto region, Gatuin Lake, Seibert 1529 (US). DARIEN: S slope of W peak of Cerro Tacarcuna Massif, Gentry & Moni 13965, 13965A (both MO). PANANIA: Hills above Campana, Allen 1691 (MO). 5 mi above Panamerican Highway on road to Cerro Azul, Croat 11521 (MO). Along trail between Rio Maje and Quebrada Brava, Croat 34636 (MO). Capira, NW side of Cerro Trinidad, Foster 2094 (DUKE). Cerro Campana, Gentry 4909 (MO). El Llano-Carti Road, 9.6 km from Panamerican Highway, Mori & Kal- lunki 1828 (MO). VERAGUAS: Rio Segundo Brasso, 8 km beyond Escuela Agricola Alto de Piedra beyond Santa Fe, Croat 25562 (MO). Along road between Escuela Agricola Alto de Piedra above Santa Fe and Rio Dos Bocas, ca. 10 km from the Escuela, Croat 25899 (MO). Valley of Rio Dos Bocas between Escuela Agricola and Calovebora, 15.6 km NW of Santa Fe, Croat 27765 (MO). NW of Santa Fe, 8.8 km from Escuela Agricola Alto de Piedra, Moni et al. 4016 (MO, USF). NW of Santa Fe, 2 km Escuela Agricola Alto de Piedra, lower slopes of Cerro Tute, Mori & Kallunki 5228 (MO, USF).
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