9. Carex scabrata Schwein.
Pl. 32 i–l; Map 125
Plants monoecious, with long-creeping
rhizomes, forming dense clumps or colonies. Vegetative growth consisting of basal
leaves with overlapping sheaths that are often broader than those of flowering
stems and usually somewhat corrugated in cross-section, located at the tips of
rhizome branches. Flowering stems 30–90 cm long, erect, sharply trigonous,
glabrous. Leaves basal and on the basal third of the stems, shorter than to
about as long as the stems, glabrous, the basal leaves with somewhat reduced
blades, the basal leaf sheaths of the previous year persistent, brown. Leaf
blades 10–40 cm long, 4–14 mm wide, thin and usually spreading to arched, the
margins and midrib minutely roughened or toothed, the margins flat. Leaf
sheaths with the tip concave, the ligule longer than wide and V-shaped, the
ventral side usually thin, papery, and yellowish white, the lowermost sheath
bases often tinged light brown. Spikes 4–9 per stem, the bracts leaflike,
mostly longer than the inflorescence, lacking a sheath or nearly so. Terminal
spike staminate, 15–40 mm long, short-stalked, linear, the staminate scales
3.5–6.0 mm long, narrowly elliptic or lanceolate, white to tan, with a green
midrib. Lateral spikes 3–8, pistillate, loosely spaced near the tip of the
axis, the uppermost sessile or nearly so, the lowermost with a long, minutely
roughened stalk, ascending, 10–50 mm long, 5–8 mm wide, narrowly oblong in
outline, with numerous densely spaced perigynia, the pistillate scales 2.0–5.5
mm long, lanceolate, pointed at the tip, light brown to reddish brown, the
midrib usually green. Perigynia 3.0–4.5 mm long, bluntly trigonous in cross-section,
the main body elliptic-obovate in outline, tapered abruptly to a beak at the
tip, tapered to a short, stalklike base, the surface usually with 2 prominent,
longitudinal ribs on opposite sides, otherwise few-nerved, roughened with
minute, stiff hairs, green, the beak 1–2 mm long, about as long as the main
body, not flattened, somewhat outwardly curved, with short, soft, inconspicuous
teeth at the tip. Styles withering during fruit development, jointed to the
main body of the fruit, which is beakless or short-beaked at maturity. Stigmas
3. Fruits 1.3–1.7 mm long, obovate in outline, sharply trigonous in
cross-section with concave sides and thickened angles, brown. May–July.
Known from a single historical collection
from Greene County (northeastern U.S. and adjacent Canada south to Georgia and west to Minnesota and Missouri). Habitat in Missouri unknown, but it should be expected
in bottomland forests and on stream banks.