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(Last Modified On 9/25/2013)
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(Last Modified On 9/25/2013)
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Species
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Centrosema pubescens Benth.
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PlaceOfPublication
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Ann. Wiener Mus. Naturgesch. 2: 119. 1837.
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Note
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TYPE: Mexico, Keerle, Herb. Martius (?M, not seen).
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Synonym
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Bradburya pubescens (Benth.) Kuntze, Rev. Gen. 1: 164. 1891.
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Description
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Twining vine scrambling on herbs and shrubs, stems slender, often wiry, puberulent to pilose. Leaves pinnate trifoliolate; leaflets ovate, apically obtuse or acute, mucronulate, basally obtuse or rounded, mostly 4-8 cm long, 1.5-3.5 cm wide, the terminal leaflet largest, pubescent with short weak erect hairs, more so beneath; petiolules tomentose, 1-3 mm long; petioles shorter than the terminal leaflet; stipels subulate, 1-2 mm long; stipules deltoid acuminate, drying striate, 1-2 mm long. Inflorescences axillary, congested several flowered racemes; pe- duncle 2-5 cm long, bracts narrow, nervate, caducous; bracteoles 2, spathaceous, 5-8 mm long, conspicuously nervate, enveloping the lower calyx teeth; pedicels ca. 5 mm long, glabrate. Flowers showy, blue or violet, almost white; calyx cupular, the lower tooth mostly subulate, often pubescent, 2-3 times as long as the others, the other teeth deltoid, 2-3 mm long, glabrate; standard 3-5 cm broad, orbicular, nearly flat, emarginate, puberulent outside, with fine dark lines and often with a purplish spot. Legume linear oblong, greatly compressed, 6-7 mm across, the margins thickened, mostly straight, 5-7 cm long, the beak prominent, slender, 0.5-1.5 cm long; seeds 15-30, compressed rectangular, dark brown, mot- tled, ca. 3 mm long.
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Habit
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Twining vine scrambling on herbs and shrubs
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Note
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This species is distinct in its calyx which has only 1 elongate, subulate tooth, the others being short and the calyx margin nearly undulate.
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Distribution
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ranges from the-southern United States to Argentina and is found in the Antilles.
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Note
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Two Panamanian collections, e.g. Folsom et al. 7174 and Lewis et al. 677, are conspicuously different in appearance from the mass of collections of this species from most of its range. The difference consists of much greater pubes- cence on bracteoles and bracts, the hairs being much denser and longer than in usual material. A number of other collections like these have been taken from widely scattered plants in Central America, but intermediates between this pu- bescent material and the usual pubescence have not been seen, suggesting that two different taxons are present. Bentham's original description of Centrosema pubescens notes bracts densely sericeous villous and bracteoles sericeous, which seems to apply more to the pubescent plants than the usual less pubescent plants, so recognition of more than one taxon might require renaming of the commoner plants rather than the unusual plants. A study of variation within the full range of material and study of the type collection will be necessary to clarify the taxons and names to be recognized here.
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Specimen
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BOCAS DEL TORO: Almirante, Cooper 76, 189 (both F). Changuinola to 5 mi S of junction of Rio Terebe, 100-200 ft, Lewis et al. 931 (MO). Water Valley, Wedel 1813 (GH, MO, US). CANAL ZONE: BARRO COLORADO ISLAND: Croat 4398 (MO), 4947 (SCZ), 6986 (F, MO, SCZ), 7031 (MO); Shattuck 715 (MO); Wetmore & Abbe 157 (F, GH); Woodworth & Vestal 500 (F, GH, MO), 716 (F). Pipeline road N of Gamboa, Correa & Haines 539 (SCZ). Monkey Hill, Cowell 45 (NY). Matachin, Cowell 197 (NY). Pipeline road, 2.3 mi from Gamboa Gate, Croat 12737 (MO). Miraflores, Croat 12860 (F, SCZ). Road C21 3 mi from Gaillard Highway, Croat 13002 (MO). Upper Rio Pequeni between Sal- amanca Hydrographic Station and Rio Boqueron, 70 m, Dodge & Allen 17280 (GH, MO). Banks of Quebrada La Palma and Cafion of Rio Chagres, Dodge & Allen 17344 (MO). France Field, Dwyer 2893 (MO). Cocoli, road to Contractors' Hill, Dwyer 7218 (MO, NY). France Field, Dwyer 8520 (F). Chagres, Fendler 70 (F, US). Anc6n Hill, Frost 2 (F). 0.4 mi SE of junction road K-2 & K-15, Graham 318 (GH). Anc6n, Greenman & Greenman 5075 (MO). Hills E of Curundii, Harvey 5194 (F). Paraiso, Hayes 112, 797 (both NY). Gattin, Hayes 815 (NY). Pipeline road, Hladik 51 (NY). Gamboa, Kennedy & Steiner 2467 (F). Pedro Miguel, Killip 3348 (US). Anc6n, Macbride & Feath- erstone 17 (F, US). Boy Scout camp near Madden Lake & dam, McDaniel 12657 (FSU). Chiva-Chiva Trail, Red Tank to Pueblo Nuevo, Piper 5144 (US). Empire to Mandinga, Piper 5145, 5156 (both US). Juan Mina, Piper 5158 (US). Anc6n, Piper 5177 (US). Chiva-Chiva Trail, Red Tank to Pueblo Nuevo, Piper 5179 (F, US). Gatuncillo, Piper 5180 (US). Old Fort San Lorenzo, Piper 5979 (US). Rio Grande near Culebra, Pittier 2078 (NY, US). Anc6n, Pittier 2763 (GH, NY, US). Balboa, Stan- dley 25270, 25486 (both US). Cerro Gordo near Culebra, Standley 25959 (US). Balboa, Standley 26067 (US). Corozal, Standley 27391 (US). Gamboa, Standley 28513, 29323 (both US). Rio Pedro Miguel near East Paraiso, Standley 29931 (US). Summit, Standley 30111 (US). Between France Field and Catival, Standley 30305 (US). Balboa, Standley 30887 (US). Dari&n Station, Standley 31576 (US). Howard Air Force Base near Red Devil drop zone, Tyson 1841 (FSU, MO, SCZ), 1846 (SCZ). 1 mi N of Summit, Tyson et al. 2743 (GH, MO, SCZ). Miraflores Locks area, Tyson & Dwyer 2741 (SCZ). Without other locality, without collector (MO-1090658). CHIRIQUI: David/Concepci6n, 60 m, Burt & Koster 135 (MO). Near Frontera, 100-200 m, Burt & Koster 147 (MO). Volcakn/Cuesta de Piedras, 1000-1200 m, Burt & Koster 158 (MO). Puerto Armuelles, Busey 513 (MO). Quebrada Cafia Blanca, Burica Peninsula, 40 m, Croat 22578 (MO). Baja Gualaca, 0-100 m, Koster 171 (MO). Quebrada Guanabanito beyond La Represa 2 mi SW of Puerto Armuelles 0-200 m Liesner 129 (NY). Quebrada Punta de Piedra, 2 mi SW of Puerto Armuelles, 0-100 m, Liesner 346 (MO). Rio Dupi, Pittier 5241 (NY, US). COCLE: El Valle de Ant6n, Burt & Rattray 39 (MO); Croat 13290 (MO); Dwyer 11957 (MO, US); Lewis et al. 2566 (MO). Aguadulce, Pittier 4954 (US). 10 mi E of Nata at Rio Grande, Tyson 5216 (FSU, MO, SCZ), 5250 (FSU, SCZ). 4 mi W of Anton on Rio Chico, Tyson & Blum 2596 (MO). Penonom&, 50-100 ft, Williams 121 (NY). COLON: 10 mi SW of Portobelo, 2-4 mi from coast, Liesner 1091 (MO). DARIEN: Between Pinogana and Yavisa, 15 m, Allen 271 (GH, MO, US). Manen&, Kirkbride & Bristan 1566 (NY). Yaviza along Rio Chucunaque, Stern et al. 150 (GH, US). HERRERA: Chitr&, 20 m, Allen 1089 (GH, MO, US). Ocui, 100 m, Allen 4088 (MO). LOS SANTOS: 1 mi N of Tonosi, Tyson et al. 2992 (MO, SCZ). PANAMA: Taboga Island, Allen 136 (GH). La Chorrera/Bejuco, 0-100 m, Burt & Rattray 24 (MO). Punta Chame, D'Arcy 10235 (MO). Quebrada La Palma and Canon of Rio Chagres, Dodge & Allen 17344 (GH). Taboga Island, Dwyer 2096 (MO). Rio Coronado along Panamerican Highway, Gentry 2906 (MO). Panama, Hayes 172 (GH). Taboga Island, Hunter & Allen 136 (GH, MO). San Jose Island, Johnston 888, 1010, 1062 (GH). Chiman, Lewis et al. 3354 (MO). Bellevista, Macbride 2741 (F). Nuevo Emperador, Munioz 35 (MO). La Cresta, Sandoval 13 (MO). Big swamp E of Rio Tocumen, Standley 26639 (US). Corozal road near Panama, Standley 26809 (US). Taboga Island, Standley 27905 (US). Rio Tapia, Standley 28191 (US). Tumba Muerto Road near Panama, Standley 29703, 29752 (US). Between Las Sabanas and Matias Hernandez, Standley 31817 (US). SAN BLAS: Mouth of Ailigandi river to 2.5 mi inland, Lewis et al. 76 (GH, MO, US); Warner 178 (MO). VERAGUAS: Divisa/Santiago, 0-100 m, Burt & Rattray 74 (MO). Puerto Mutis 12 mi S of Santiago, Tyson 5190 (FSU, MO, SCZ), 6007 (SCZ).
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