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(Last Modified On 10/15/2013)
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(Last Modified On 10/15/2013)
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Species
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Spachea membranacea Cuatr.
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PlaceOfPublication
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Webbia 13: 548-549. 1958.
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Note
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TYPE: Rio Carepa, Quebrada Entre-Dos, 12 km NE of Chigorodo, 35 km S of Turbo, 100 m, Antioquia, Colombia, Haught 4732 (US, holotype).
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Description
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Slender trees, to 10 m tall; sap often somewhat milky; young stems, petioles, inflorescence branches, and midrib of leaf strigose with ferruginous, T-shaped trichomes. Leaves opposite, ovate-elliptic to oblanceolate, long-acuminate, rounded or shortly. cuneate at the base, 4-18 cm long, 1.5-6.0 cm wide, glabrate except for the midrib above and below, with several round glands below, usually 1 on each side of the midrib below and others sometimes scattered throughout the blade, the margin entire, pubescent, revolute in age; petioles 6-9 mm long; stipules connate, minute, ovate, axillary. Inflorescences of long, pendent, axillary racemes 10-20 cm long, longer than the leaves, the axes and peduncles sparsely ferruginous-pubescent with appressed, T-shaped trichomes; axis slender, flex- uous; bracts ovate-oblong, acute, ca. 1.5 mm long, united to the peduncle at the base; pedicel 1-flowered, 2-5 mm long, bearing 2 small bracts midway and 5 inconspicuous glandular appendages at the apex; peduncle 5-10 mm long, pink. Flowers 8 mm diam.; calyx 8-glandular, the sepals oblong-elliptic, ca. 2.5 mm long, thickened toward the apex, puberulent or glabrescent, acute to rounded and reflexed at the apex, ciliate, the glands white, prominent, 2.5-3.0 mm long, nar- rowed and curved outward at the apex; petals clawed, magenta, obovate, ca. 4 mm long, ciliate, spreading, the base attenuate, the margins weakly denticulate, the claw thick, 1.0-1.5 mm long; stamens 10, ca. 2 mm long, shorter than the sepals, subequal, the anthers elliptic-oblong, glabrous, ca. 1 mm long, the fila- ments ca. 1 mm long, hirsute at the base; pistil 2-parted, each ovary with a slender, persistent, divergent style, longer than the ovary, the apex truncate, thickened, held at about the level of the anther. Fruit a coriaceous schizocarp of 2 separable, 1-seeded carpels, each carpel indehiscent, ovoid, 3.5-5.0 mm long and ca. 2 mm wide, with a thin fleshy red covering at maturity; seeds hemi- spheroid, tan, smooth, ca. 4 mm long.
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Habit
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trees
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Distribution
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known from Colombia and Panama.
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Note
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In Panama the species occurs in tropical moist forest at less than 500 meters elevation. It is perhaps closest to S. herbert-smithii (Rusby) Cuatr. from Colombia, but that species has a prominent stalked gland on each 1-flowered peduncle.
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Specimen
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CANAL ZONE: Upper Chilibre River, 1 mi below Chilibre, Seibert 1511 (MO, US). BARRO COL- ORADO ISLAND: Aviles 12 (F, MO), 1007 (F); Bangham 579 (A, F); Croat 4748 (MO, NY, SCZ), 4756, 4769, 4778 (all MO, SCZ), 6397 (GH, MO, SCZ), 8123, 8470 (both MO), 15150 (DUKE, GH, MO, NY, SCZ), 14625 (MO), 14862 (DUKE, F, MO, NY, PMA, SCZ), 14920 (F, MO, SCZ), 16632 (DUKE, GH, MO, NY); Ebinger 172 (MO, US); Foster 873, 1055, 1078 (all DUKE, F, GH, MO, PMA), 1342 (DUKE); Knight 4035 (MO); Salvoza 1000 (A); Shattuck 1007 (MO); Webster & Breckon 16441 (DUKE, MO). DARIEN: Rio Perecenico, Duke 8221 (MO, OS). Quebrada Maskia off Rio Pucro above Pucro, Duke 13092 (MO, OS, US). El Real, Rio Tuira, Stern et al. 767 (MO). Campamente Buenevista, Rio Chucunaque above confluence with Rio Tucesa, Stern et al. 922 (MO). PANAMA: Between Rio Maje and Quebrada Brava, 60 m, Croat 34607 (MO). Rio Maje, ca. 4-5 mi above waterfalls near Bayano Lake, 100 m, Croat 34694 (DUKE, MO, NY, RSA, US). Rio Potrero, ca. 2 mi from Arrajan, Duke 4683 (G, MO, NY). Between Cafiasas and Sabola, ca. 100 m, Duke 14459 (MO, OS, US). Bayano, Ciupo forest, Rio Piarti, Folsom 3543 (MO). SAN BLAS: Hydro Camp Cuadi on Rio Cuadi, Duke 15460 (US).
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