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(Last Modified On 1/14/2013)
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(Last Modified On 1/14/2013)
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Species
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COCCOLOBA MANZANILLENSIS Beurling
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PlaceOfPublication
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Prim. Fl. Portobello in Kongl. Vetensk. Akad. Handl. Stockh. 142. 1854 (1856).
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Reference
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Lindau, Engl. Bot. Jahrb. 12:209. 1890.
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Synonym
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Campderia nematostachya Griseb., Bonplandia 6:4. 1858. Coccoloba nematostachya Lindau, Engl. Bot. Jahrb. 13:208. 1890.
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Description
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Tree; branches striate, canescent, branchlets striate, silky pubescent; ocreae membranaceous to thinly coriaceous, divided to the middle, protracted in a long obtuse apex, to 4.5 cm. long, golden or tawny pubescent; petioles inserted at the base of the ocreae, 1-2 cm. long, canescent on the adaxial surface, short pilose elsewhere becoming glabrate; leaf blades oblong-elliptic, obovate or obovate-elliptic, apex short acuminate to rarely rounded, narrowed below the middle to a cordate or nearly rounded base, 11 X 6, 15 X 8, to 25 X 15 cm. long and wide, thin, cori- aceous and flat when young, conspicuously undulate or bullate between the veins when mature, margin reflexed, primary veins 14-17 on each side, generally glabrous but long pilose on the midrib; inflorescence terminal, the staminate 15-34 cm. long, the pistillate 20 cm. long, loosely flowered, all parts pilose, peduncle 1 cm. long, staminate flowers in clusters of 3 or 4, pistillate flowers borne singly at each nodule, bracts and ocreolae minute, to 0.5 cm. long, pilose, pedicels thin, about 1 mm. long, hypanthium campanulate, slightly stalked, 1 mm. long, the perianth lobes ovate, about 1 mm. long, fertile filaments 1.5 mm. long, exserted, fertile pistil 1.5 mm. long, styles 3, fruiting pedicels 1-1.5 mm. long, fruit ovoid, slightly stalked at the base, coronate at the apex, strongly striate when dry, 7 mm. long, 4 mm. thickest diameter; achene dark brown, strongly trigonous.
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Habit
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Tree
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Note
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Local name: "'Hueso" (ex Standley). Collected in flower in January. Collected in fruit in June.
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Common
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Hueso
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Native
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Endemic to Panama
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Distribution
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Panama
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Elevation
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10-20 m.
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Specimen
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CANAL ZONE: Barro Colorado Island, L. H. d E. Z. Bailey 42; Shattuck 425; between Gatun and Lion Hill, Pittier 2570. COLON: Porto Bello, Billberg 230, 234 (Holotype). DARIEN: La Palma, Pittier 6599. PANAMA: Pacora, Allen 3450. PROVINCE UNKNOWN: Andersson s. n. (Type of C. nematostachya Griseb., GOET.)
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Note
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The appearance of specimens assigned to this species vary considerably and depend on the period of collection. The leaves are apparently deciduous and young foliage appears with the flowers. Thus flowering specimens have smaller leaves of soft texture which turn black on drying. This is the condition of the type speci- mens of both species considered here. A few leaves of the Andersson collection, which is the type of C. nemafostachya, are more mature and show the bullate characteristic also found in the fully mature leaves of the collection by Liberty Hyde and Ethel Zoe Bailey (number 42) which is the only collection known in fruit. The long ocreae and copious silky pubescence clearly mark this species. The original spelling of C. manzinellensis by Beurling has been changed to C. manzanillensis by Lindau. While the island where Billberg collected the type speci- men can not be located on modern maps, Beurling's intentions were obvious. Manzanillo, or "little apple" in Spanish, is commonly applied to the widespread and infamous strand plant Hippomane manchineel. Although Lindau failed to com- ment on his correction of an orthographic error or to cite the original spelling, it seems desirable to allow this correction to stand.
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