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(Last Modified On 1/14/2013)
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(Last Modified On 1/14/2013)
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Species
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COCCOLOBA OBOVATA HBK.
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PlaceOfPublication
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Nov. Gen. 2:176. 1817.
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Synonym
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Coccoloba Goudotiana Wedd. Ann. Sci. Nat. III, 13:260. 1850. Coccoloba riparia Lundell, Contr. Univ. Mich. Herb. 6:11. 1941.
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Description
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Shrub of 3 m. to tree 15 m. tall; branches terete, glabrous; ocreae oblique, truncate, 7-10 mm. long, glabrous, membranaceous, striate, appressed; leaves of normal shoots with petioles 12-15 mm. long, pilose or glabrous, inserted above the base of the ocreae, blades obovate to subround-obovate, rarely ovate-lanceolate to elliptic obovate, 9 X 6, 10 X 4, 16 X 8 to 24 X 9.5 cm. long and wide, apex obtusely acuminate, base slightly cordate, coriaceous, turning dark brown or black on drying, tomentulose on the midrib and veins, persistent, short pilose in the axils of the veins and midrib, commonly densely covered with globose or peltate excretions associated with the stomata; leaves of adventitious shoots with petioles 3.5-6 cm. long, blades broadly elliptic-ovate to elliptic-oblong, 28 X 14 to 37 X 26 cm. long and wide; inflorescence terminal, solitary, densely flowered, staminate to 25 cm. long, pistillate to 17 cm. long, the flowers borne singly in the pistillate and in clusters of 4 in the staminate, peduncle, rachis, bracts and ocreolae short pilose, bracts semi-orbicular, 0.75 mm. long, ocreolae bibbed 1.5 mm. long, mem- branaceous, usually in multiples, pedicels included in the ocreolae in flower, exserted and to 1.5 mm. long in fruit, hypanthium 0.75 mm. long, the lobes ovate or nearly orbicular, to 1 mm. long, functional stamens exserted, sterile stamens included, ovary triangular, styles 3; fruit globose, obtusely-trigonous, strongly striate, the base rounded, the apex obtusely coronate, 6 mm. long, 5.5 mm. in diameter; achene tan to dark brown, trigonous.
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Habit
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Shrub tree
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Note
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Type: Colombia, prov. Tolima, Honda, Humboldt s. n. June 1805 (Herb. Willd. 7704), isotype Paris. Common name: "Papaturra blanca", "Pefia blanca".
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Common
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Papaturra blanca Pefia blanca
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Note
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Collected in flower in March, June, July, September and December. Collected in fruit in August and December.
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Elevation
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50 to 800 m.
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Distribution
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Costa Rica, Panama, Colombia.
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Specimen
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CANAL ZONE: Barro Colorado Island, Aviles 998, Salvoza 984, Shattuck io63, Starry 298; F. L. Island, Bangham 593; Salamanca Hydrographic Station, Dodge, Steyermark d Allen i6983, i6983a, Woodson, Allen d Seibert i572; northwestern part of Canal Zone, Johnston I5i9; between Tumba Vieja and Salamanca, Steyermark d Allen i6758. CHIRI- QuI: Cerro Galera Chorcha, Gualaca, Allen 5020; San Felix, Allen 3655; between San Felix and Cerro Flor, Allen I9I7; Progreso, Cooper d Slater 270; San Bertolome, Woodson d Schery 947; between Rio Chiriqui and Remedios, Woodson, Allen d Seibert ii82. COCLE: El Valle, Allen 2229 (MICH-holotype of C. riparia). VERAGUAS: between Santiago and David, Stern, Chambers & Dwyer and Ebinger 999.
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