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(Last Modified On 6/6/2013)
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(Last Modified On 6/6/2013)
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Species
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Neomirandea homogama (Hieron.) H. Robinson & R. D. Brettell
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PlaceOfPublication
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Phyto- logia 28: 62. 1974.
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Synonym
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Liabum homogamum Hieron., Bot. Jahrb. Syst. 28: 626. 1901. TYPE: Colombia, near La Conga, region Popayan, Lehmann 5972 (F, photo and fragment, US). Eupatorium hitchcockii B. L. Robinson, Contr. Gray Herb. 73: 14. 1924. TYPE: Ecuador, valley of Pastaza River between Bafios and Cashurco, Tungurahua, Hitchcock 21876 (US). Neomirandea hitchcockii (B. L. Robinson) R. M. King & H. Robinson, Phytologia 19: 308. 1970.
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Description
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Coarse herbs or shrubs to 5 m tall, sometimes epiphytic; stems terete or grooved, subglabrous, fistulose. Leaves opposite; blades papyraceous to slightly fleshy, ovate to oblong-ovate, to 30 cm long and 20 cm wide, the base cuneate to truncate, the margins serrate to slightly dentate, the apex sharply acute to slightly short-acuminate, the surface glabrate, the venation pinnate with many pairs of secondaries often concentrated in basal fourth; petioles 4-6 cm long, glabrate to sparsely puberulous. Inflorescence a loose corymbose panicle, to 20 cm wide and 20 cm high, the ultimate branches 3-12 mm long, densely puberulous. Heads 9-11 mm high with 14-21 florets; involucral bracts ca. 20-25, imbricate, in 3-4 series, mostly 2-6 mm long, the apices obtusely acute to narrowly rounded, the outer surface minutely puberulous to glabrate, the outer bracts broadly ovate to sub- orbicular, the inner bracts broadly oblong to oblong-lanceolate; receptacle puberulous; corolla lavender to purple, 5-7 mm long, funnelform with narrow tube as long as or longer than limb, with numerous hairs on inner surface, few hairs on outer surface, the lobes 0.8-0.9 mm long aiid 0.6 mm wide, with few nearly sessile glands and few short hairs on outer surface; style base enlarged, glabrous. Achenes prismatic, 3-4 mm long, glabrous; carpopodium distinct, with small cells in 8-10 series; pappus of ca. 40 slightly scabrous bristles, mostly 5-7 mm long, the apical cells acute.
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Habit
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herbs or shrubs
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Distribution
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known from Panama southward through Colombia to southern Ecuador.
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Note
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The species is one of a pair that have previously been dis- tinguished primarily by geography and number of florets per head. The presence of both species in Panama and the greater variation in number of florets have resulted in some problems in delimiting the species, and the name N. homogama was at one point applied to the related N. standleyi. The two species are now recognized as very close relatives with nearly continuous if not slightly over- lapping ranges. The more eastern N. homogama has larger heads with more prominent, often broadly ovate, outer involucral bracts and 14-21 florets per head. The corollas are generally narrower with tubes usually as long as the limb. The more western N. standleyi has smaller heads with usually oblong or elliptical outer involucral bracts that do not project as strongly on their basal margins. There are 5-12 florets per head and the corollas are more generally flaring with tubes distinctly shorter than the limbs. The distribution of characters suggests that two thoroughly distinct species exist with possibly some complications through hybridization.
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Specimen
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BOCAS DEL TORO: Chiriquf trail between Quebrada Higueron and Gutierrez, Kirkbride & Duke 750 (MO, REED, US). COCLE': La Mesa above El Valle, Croat 14304 (MO).
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