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(Last Modified On 7/16/2013)
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(Last Modified On 7/16/2013)
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Species
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Paullinia fasiculata Radlk.
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PlaceOfPublication
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Monogr. Paull. 124. 1895-1896.
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Note
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TYPE: Guiana, Poiteau (G).
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Description
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Lianas; stems sharply triangular when young, glabrate, becoming markedly 3-ribbed in age and sometimes appearing as 3 separate strands only loosely at- tached; wood composite with an irregular central core and three smaller periph- eral bundles, these separating in age, the periderm coarse, somewhat exfoliating in age. Leaves pinnately 3-5-foliolate; petioles 3-13 cm long, narrowly winged, glabrous or scabridulous at the apex and sparsely so along the margins of the wings; rachis of the 5-foliolate leaves winged, the wing similar to the petiole wing; leaflets elliptic to ovate- or oblong-elliptic, gradually to abruptly acuminate to acute at the apex, acute to obtuse at the base, 7-20 cm long, 3-9 cm wide, glabrous or puberulent only on the midrib above, glabrous beneath except for some barbul- ate vein axils, the margins sparsely scabrid, the reticulate veins prominulous on both surfaces; juvenile leaves simple, 15-25 cm long, 8-14 cm wide; stipules linear- lanceolate, 3-4 mm long, striate, glabrous inside, appressed-pubescent outside, often deciduous. Inflorescences of densely glomerulate thyrses borne on raised protuberances from the central strand of the stem; rachis branched, to ca. 2 cm long; rachis, pedicels, and calyces sparsely appressed-puberulent to glabrate, dry- ing dark brown; bracteoles narrowly triangular, less than 1 mm long; pedicels to ca. 6 mm long. Flowers sweetly fragrant, white; buds broadly oblong, somewhat flattened laterally; outer sepals 1.5-2 mm long, ellipsoid, sparsely appressed- pubescent outside, glabrous within, the inner sepals obovate, appressed-puberu- lent outside, glabrous within; petals oblong-obovate, 3-4.5 mm long, the scales 2/3_4 as long as the petals, the basal part rounded, 1.5 mm long, glabrous or with the margin sparsely ciliate, the crest obovate to inequilaterally emarginate, thin, orange, the deflexed appendage shorter than or equalling the crest, rounded at the base, glabrous or densely villous; disc glands white, semicircular or longer than broad, puberulent to appressed-puberulent in at least the lower half within; stamens with the filaments flattened, loosely villous in the lower half; staminate flowers with the stamens basally united, to ca. 3 mm long, the pistil minute, ap- pressed-pubescent on the upper half of the ovary and on the style and stigmas; bisexual flowers with the stamens ca. 1.8 mm long, free, the ovary and style to- gether 3.5 mm long, the ovary 1.5 mm long, densely pubescent, the style 3-branched in the upper A/i, sparsely pubescent. Capsules obovoid-pyriform, long-stipitate, 1.5-2 cm long, apiculate and obscurely 3-winged at the apex, the outer surface sparsely short-pubescent toward the apex, glabrous toward the base, longitudinally striate, the inner surface apparently glabrous.
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Habit
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Lianas
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Note
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The species is distinguished by the (3-) 5-foliate leaves with winged petioles and rachises, glomerulate inflorescences with small acute bracts, rachises much longer than pedicels, and 3-sulcate stems with composite wood. In Panama flowering occurs from December to February with fruits develop- ing during April and May.
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Distribution
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ranges from Costa Rica to Panama. It is also known from Guyana and Tobago (West Indies). In Panama it occurs principally in premon- tane wet forest and tropical wet forest but also in wetter parts of tropical moist forest on the Atlantic slope.
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Specimen
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BOCAS DEL TORO: Daytonia region of Almirante, Cooper 135 (F). COLON: Lower Rio Guanche, Dressler 4238 (MO). S of Portobelo, Foster 2065 (DUKE). Rio Guanche, Foster 2198 (DUKE); Mori & Kallunki 3020 (MO, PMA); 0-10 m, Nee & Gentry 8693 (CAS, F, MO, PMA, RSA, US). PANAMA: El Llano-Carti Road, 8-12 km N of El Llano, 400-450 m, Nee et al. 8767 (F, MO, PMA, US).
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