1. Pedicularis canadensis L. (common lousewort, wood betony)
P. canadensis var. dobbsii Fernald
Pl. 472 a, b;
Map 2164
Plants often
colonial, with short rhizomes or stolons. Stems 10–30(–40) cm long, sparsely
hairy toward the base, densely hairy or appearing woolly toward the tip and
inflorescence axis. Basal leaves well-developed at flowering, long-petiolate.
Stem leaves mostly alternate. Lower stem leaves long-petiolate. Leaf blades 2–9(–15)
cm long, lanceolate to narrowly oblong-elliptic or occasionally oblanceolate,
moderately to deeply lobed, those of the larger leaves lobed more than halfway
to the midvein, the lobes finely toothed or scalloped to lobed again, usually
with a narrow, thickened pale band along the margins, the surfaces sparsely
hairy to nearly glabrous. Inflorescences terminal spikes or spikelike racemes,
the flower stalks absent or to 3 mm long, relatively short and dense when the
first flowers open, continuing to elongate and appearing less dense as the
fruits develop. Calyces 7–9 mm long, strongly oblique, the tube with a pair of
shallow, rounded teeth at the tip of the upper side (sometimes appearing merely
shallowly notched), the lower side deeply notched or divided. Corollas 18–25 mm
long, yellow or variously tinged with pink, red, or purple, rarely white, the
upper lip 11–15 mm long, sometimes completely purple to dark purple, narrowed
to a more or less truncate tip, but with a pair of short, slender, toothlike lobes
just below the tip, the lower lip 7–10 mm long, the lobes spreading, the
central lobe shorter and more sharply pointed than the lateral ones. Fruits
10–17 mm long. Seeds 1.6–2.2 mm long, more or less oblong-ellipsoid, not
flattened, with a short, sharp point at the tip, truncate to concave at the
base, unwinged, the surface with a fine but noticeable network of ridges,
brown. 2n=16. April–May.
Scattered nearly
throughout the state, less abundantly north of the Missouri River (eastern U.S.
west to North Dakota and Texas; Canada). Glades, upland prairies, ledges and
tops of bluffs, savannas, openings of mesic to dry upland forests, and less
commonly banks of streams and rivers and openings of bottomland forests; also
old fields, pastures, and roadsides.
Viewed from
above, the inflorescences of this species often appear similar to pinwheels,
with the flowers strongly spiraled. Flower color can vary greatly within
populations. Rare plants with the corollas white or mostly white have been
called f. albescens Steyerm. Steyermark (1963) noted anecdotal reports
that P. canadensis is toxic to sheep.