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(Last Modified On 5/9/2022)
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Acceptance
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Accepted
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(Last Modified On 7/18/2018)
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Contributor Text:
R.R. Mill
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Contributor Institution:
Royal Botanic Garden, Edinburgh, Scotland
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Synonym Text:
P. tubiformis Klotzsch, Bot. Ergebn. Reise Waldemar 106, t. 57. 1862; P. longiflora Rudolph var. tubiformis (Klotzsch) P.C. Tsoong, Acta Phytotax. Sin. 3: 278 & 318. Jan. 1955; Polunin & Stainton, Fls. Himalaya 300. 1984; Hanbi et al. in Wu Zheng-yi & P. R. Raven Fl. China 18: 205. 1998.
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Flower/Fruit:
Fl. Per.: July – October.
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Type:
Lectotype: the illustration of Pedicularis tubiformis in Klotzsch & Garcke, Bot. Ergebn. Reise Prinz Waldemar Preuss. : t. 57. 1862.
“Dr Hoffmeister sammelte diese Art im Himalaya.” Type: NW India (Garhwal to Bashahr), 1846, W. Hoffmeister s.n. Records of collecting localities were lost due to Hoffmeister’s violent death in the field (Pennell l.c.). Hoffmeister's types were at B (now destroyed; no duplicate at C, database searched 26.01.01).
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Distribution:
Pakistan, Kashmir, NW India, Nepal, Sikkim, Bhutan, China.
Springs, seepage areas, along streams and in open wet alpine meadows, marshland and irrigated fields; 3800–5200 m.
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Comment/Acknowledgements:
The flower is used in Ladakh for lung and liver diseases (Brown & Rothera 132 in sched.).
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Map Location:
B-8 or B-9 Baltistan: Kasurmik, 9000 ft., 15.8.1940, R. R. Stewart 20827 (RAW; GH–n.v.); B-9 Baltistan: Thalle La, 10,000 ft., yellow, 13.8.1940, R. R. Stewart 20619 (RAW); Ladakh: Hanle, Rupshu, in wet meadow, 14,300 ft., flowers 13.7.1931, W. Koelz 2303 (K, RAW); Kanji, 12,900 ft., fertile field (alp) by riverside, 5.7.1984, A. R. Brown & M. A. Rothera 132 (K); Khardong La, Leh, Karakoram, 15,000 ft., marsh ground, 19.8.1928, Ludlow 492 (BM); Khardong Pass, 15–17,000 ft., 7.1937, C. C. Burt 156 (E); Ladak, Gya, Watt 2435 (E); Leh, Ladakh, 11,500 ft., marsh land, 8.7.1941, Ludlow & Sherriff 8475 (BM, E); Leh to Norla, right side of the Indus valley, 15.9.–5.10.1856, Schlagintweit Cat. 1595 (E); Phola La Pass, 12,500 ft., 27.7.1937, C. C. Burt 66 (E); Shushak, 14,300 ft., grassy banks of streams, 1.8.1931, Ludlow 833 (BM); Stok, irrigated and cultivated land around village, 3352 m, 1976, Janet Maxwell 1929 (E); Tangyar Shyok valley, 12,600 ft., 14.8.1947, B. C. F. Schomburg 51 (BM); villages on the left side of the Indus valley, 20 miles SW of Leh, 1–25.9.1856, Schlagintweit Cat. 1794 (BM); Tibet/Ladakh: Bog near Moolbekh, 10,700 ft., July, 1894, W. W. Cordeaux 1896 (BM); Ladakh: unloc., Jaeschke 206 (K).
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Perennial herbs, 10-50 cm high with several, fleshy, elongate, fusiform roots to 8 cm. Stems 10–50 cm, usually several, simple, decumbent to erect, glabrous. Leaves alternate with the upper ones sometimes subopposite and clustered; petioles 0.5–7.5 cm long, narrowly winged with broadened, flattened base, glabrous or sparsely pilose; lamina linear to oblong, 10–40 x c 3–10 mm, pinnatifid segments 6–10 pairs, ovate or ovate-oblong, acute, doubly callose-serrate, glabrous. Inflorescence an abbreviated, several-flowered, terminal raceme; flowers opening from below or all flowers almost simultaneously, often with some axillary flowers below. Bracts leaf-like, with petioles broadened at base. Pedicels 2–15 mm long; flowers strictly erect at anthesis, strongly recurved and touching ground after pollination. Calyx urceolate-tubular, narrowed at mouth, 2-lobed, 11 × 4–5 mm, split to c. halfway on ventral side; teeth 2, shortly stipitate, ovate, 3-fid, the lobules acutely dentate. Corolla bright golden yellow, with 2 reddish-brown, chocolate or purple, comma-like marks in throat and pale yellow tube, slightly scented; tube 25–50 × 1.5 mm, narrowly cylindrical, pubescent; lower lip 3-lobed, 6–10 × 15–18 mm, middle lobe obovate-orbicular, 5–6 × 5.5–6 mm, lateral lobes reniform, c. 10 × 8 mm, emarginate, lobes pilose on margin; galea with lower part c. 4 mm long, abruptly bent at top, anther-bearing part 5–6 × 2.5–3 mm, directed forwards but becoming strongly twisted, narrowed into a slender beak c. 5 mm long with bilobed apex (turning orange-red after pollination). Stamens inserted at corolla tube apex; filaments pilose; anthers oblong, c. 2 × 1 mm, thecae acute at base. Capsule lanceolate-ellipsoid, 13–17 × 4–5 mm, acuminate.
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