Plants erect in loose to dense tufts 0.5-8 cm high, simple or seldom forked, often radiculose, mostly equally foliate. Leaves long lance-acuminate, erect or slightly curved, straight or ± flexuose; costa broad (more than 1/2 the leaf base), percurrent or excurrent, sometimes ending in a hyaline point, often ribbed and sometimes serrulate at back, with 0, 1, or 2 stereid bands; alar cells none or conspicuously differentiated, hyaline or brownish-red; basal cells enlarged, with thin or thick walls, sometimes porose, the outer cells often narrow and elongate; upper cells quadrate, short-rectangular, shortly elongate or rhomboidal. Dioicous. Perichaetial leaves broader at the base and long-pointed than stem leaves. Setae generally cygneous-curved; capsules ovoid to short-cylindric, often furrowed; operculum rostrate; peristome teeth 16, deeply divided, vertically pitted-striolate below. Calyptrae cucullate, usually fringed.