5. Eurhynchium Bruch & Schimp. in B.S.G.   美喙藓属   mei hui xian shu

Bryol. Eur. 5: 217. 1854. Hypnum subgen. Eurhynchium (Bruch & Schimp.) Hobk., Syn. Brit. Mosses 152. 1873. Hypnum sect. Eurhynchium (Bruch & Schimp.) Sull. in Gray, Manual (ed. 2) 669. 1856.

 

Plants slender to rather robust, pale green to dark green, glossy, in loose or dense mats. Stems prostrate or ascending, or sometimes dendroid, irregularly subpinnately branched; branches julaceous or complanate. Stem and branch leaves similar or differentiated; stem leaves broadly ovate, oblong-ovate, or triangularly ovate-lanceolate, concave, usually plicate, acute, shortly to long acuminate; slightly decurrent or barely decurrent at base; margins usually serrulate to serrate; costae single, reaching above mid-leaf, often projecting at tips; median and upper leaf cells linear or linear-rhomboidal, smooth; basal cells shorter, broader; alar cells differentiated, subquadrate, shortly rectangular or elliptical; branch leaves smaller or clearly larger than stem leaves, sometimes with twisted leaf apices. Dioicous or autoicous. Perichaetial leaves sheathing at base, often twisted above. Setae elongate, roughened or smooth; capsules inclined or horizontal, ovoid or elongate-ovoid; stomata present; opercula long rostrate; peristome double; exostome teeth papillose above, cross-striolate below, bordered, trabeculate on ventral surface; endostome segments mostly as long as the teeth, linear-lanceolate, keeled, perforate; basal membrane high; cilia present, nodulose or appendiculate. Calyptrae cucullate, smooth. Spores spherical, smooth or papillose.

 

1. Plants dendroid; stem and branch leaves sharply differentiated; stem leaves broadly ovate to semi-orbicular; branch leaves triangularly ovate-lanceolate, abruptly and slenderly acuminate at apex............. 2. E. arbuscula

1. Plants prostrate; stem and branch leaves slightly differentiated; stem leaves elliptical, ovate to ovate-lanceolate; branch leaves oblong-ovate or triangularly lanceolate....................................................................... 2

2. Leafy branches complanate................................................................................................................................. 3

2. Leafy branches julaceous..................................................................................................................................... 4

3. Leaves oblong-elliptical, usually acute at apex; margins serrate nearly throughout...................... 8. E. laxirete

3. Leaves ovate-lanceolate, usually narrowly acute to acuminate; margins only serrulate above............10. E. savatieri

4. Leaf apices acute.................................................................................................................................................. 5

4. Leaf apices shortly acuminate to slenderly acuminate..................................................................................... 7

5. Stems irregularly pinnately branched; median leaf cells linear-rhomboidal, 60–87 µm × 6–8 µm............ 1. E. angustirete

5. Stems regularly pinnately branched; median leaf cells elongate-rhomboidal, 35–56 µm × 4–7(–8) µm........ 6

6. Plants usually less than 2 cm long; leaves broadly ovate to elliptical, shortly acute at apex; median leaf cells 54–56 µm × 4–6 µm.................................................................................................................... 6. E. hians

6. Plants to 7 cm long; leaves oblong-ovate, narrowly acute to acuminate at apex; median leaf cells 35–50 µm × 7–8 µm........................................................................................................................... 9. E. longirameum

7. Leaves ending in a long, piliferous apex, distinctly longitudinally plicate................................... 3. E. coarctum

7. Leaves shortly or narrowly acuminate, not ending in a piliferous apex, not longitudinally plicate.................. 8

8. Plants filiform; leaves small, mostly 0.7–0.9 mm × 0.3–0.4 mm............................................. 5. E. filiforme

8. Plants not filiform; leaves large, usually 1.7–2.0 mm × 0.7–0.9 mm............................................................. 9

9. Stem leaves triangularly ovate-lanceolate, often squarrose at apex, broad at insertion; median leaf cells elongate-rhomboidal, 40–57 µm × 5.5–8.0 µm.................................................................... 11. E. squarrifolium

9. Stem leaves ovate-lanceolate or oblong-elliptical, not squarrose at apex, strongly constricted at insertion; median leaf cells linear to long linear, 60–190 µm × 5–6 µm........................................................................... 10

10. Leaves ovate-lanceolate, with straight acuminate apices; median leaf cells 60–97 µm long................ 4. E. eustegium

10. Leaves elliptical, with twisted apices; median leaf cells 130–190 µm long.................... 7. E. kirishimense