Plants medium-sized to robust, in loose or dense, green, yellow-green, or reddish-tinged tufts. Stems simple or sparsely forked, radiculose nearly to the apex, sometimes bearing terete, microphyllous branchlets in leaf axils. Leaves crisped when dry, tubulose below, keeled above, gradually to abruptly acuminate from an ovate or oblong base, acute; margins erect or slightly incurved and entire or toothed above, incurved below; costa percurrent or nearly so, 1/15-1/6 the leaf base, smooth or toothed at back near the apex, with 2 stereid bands; upper cells irregularly rounded, quadrate, or rectangular, smooth or bulging, with walls mostly thick and sometimes sinuose and pitted; basal cells rectangular to linear, the alar cells rounded to oblong, enlarged or inflated, orange or brownish, often extending to the costa. Dioicous (usually, if not always, pseudautoicous). Inner perichaetial leaves convolute-sheathing at base, often with a long, setaceous apex extending to the base of the capsule or beyond. Setae single or rarely paired, elongate, straight or slightly flexuose, smooth, yellow to brown; capsules erect and symmetric or sometimes slightly curved, cylindric to ovoid, smooth, yellow to brown; annulus none; operculum short- to long-rostrate; peristome teeth undivided and perforate or variously cleft, papillose throughout. Spores spherical to ovoid. Calyptrae cucullate, smooth, entire at base.