Tropicos | Name - !!Tortula Hedw.

Bolivia Bryophytes > Pottiaceae > Tortula

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!!Tortula Hedw.

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Project Name Data (Last Modified On 6/19/2009)

Acceptance : Accepted

Project Data (Last Modified On 10/30/2017)

Plant Category: Moss

Notes (Last Modified On 10/30/2017)

general taxon notes:

Tortula is a widespread genus of ca. 140 species; about 11 species in South America (Cano & Gallego 2008). A single species is presently recognized for Bolivia. The genus is characterized by the smaller but not thicker-walled epidermial stem layer (ca. same thickness as the immediate internal cells), presence of central strand, obovate, spathulate to oblong leaves, often recurved and entire margins, subpercurrent to excurrent costa, in cross-section costa semicircular to rounded in shape, usually with well developed surface cells, and mostly a well developed dorsal stereid band. Previously Tortula was broadly defined, containing species that have since been placed in various genera including for Bolivia: Chenia, Dolotortula, Hennediella, Sagenotortula, and Syntrichia. The following treatment and key is adapted from Cano and Gallego (2008). At least three, possibly four, additional species may be found in Bolivia.

Reference List (Last Modified On 10/30/2017)

Reference:

Cano, M. J. & M. T. Gallego. 2008. The genus Tortula (Pottiaceae, Bryophyta) in South America. Bot. J. Linn. Soc. 156: 173–220.

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1. Middle external marginal cells ofthe leaf linear to long-rectangular (length/width ratio 7.5-40); capsulehorizontal or cernuous (recorded for northwest Argentina) … T. cernua

1. Middle external marginal cells ofthe leaf quadrate, subquadrate, short-rectangular, or oblate (length/width ratio 0.6-1.9); capsules erect … 2.

2. Upper and middle laminal cellssmooth or with 1-3 simple papillae, 1 µm high (southern Peru) … T. arequipensis

2. Upper and middle laminal cellswith 4-6 simple or bifurcate papillae, 2.5-5 µm high … 3.

3. Ventral surface cells of thecosta rounded to rectangular, 22.5-25 µm in length, bulging, disposed in1-2 layers in cross-section … T.atrovirens

3. Ventral surface cells of thecosta quadrate, rectangular, sometimes subquadrate, rarely oblate or rounded 10-17.5 µm in length, not bulging, disposed in 1 layer in cross-section… 4.

3. Middle laminal cells 12.5-20 µmwide; peristome teeth ca. 500 µm long, spirally twisted (Peru?) … T. vahliana

3. Middle laminal cells 7.5-12.5 µm wide ; peristome teeth ca. 100-132 µm long (recorded for Peru) … T. platyphylla

1. Células mediales externas del margen de las hojas lineales a largo-rectangulares (proporción largo/ancho 7.5-40); cápsulas horizontales o cernuas … T. cernua

1. Células mediales externas del margen de las hojas cuadradas, subcuadradas, corto-rectangulares, u oblatas (proporción largo/ancho (0.3)0.6-1.9(2.8)); cápsulas erectas … 2.

2. Células superiores y mediales de la lámina lisas o con 1-3 papilas simples, 1 µm de alto … T. arequipensis

2. Células superiores y mediales de la lámina con (2)4-6(10) papilas simples o bifurcadas, 2.5-5 µm de alto … 3.

3. Células mediales de la lámina de12.5-20 µm de ancho; dientes del peristoma aproximadamente 500 µm de largo, torcidos en espiral … T. vahliana

3. Células mediales de la lámina (5)7.5-12.5(15) µm de ancho ; dientes del peristoma aproximadamente 100-132 µm de largo … T. platyphylla

1. Células mediales externas del margen de la hoja lineares a largo-rectangulares (proporción longitud/ancho 7.5-40); cápsula horizontal o cernuas (registrado en el noroeste de Argentina) … T. cernua

1. Células mediales externas del margen de la hoja cuadradas, subcuadradas, corto-rectangulares u oblatas (proporción longitud/ancho (0.3)0.6-1.9(2.8)); cápsulas erectas … 2.

2. Células superiores y mediales de la lámina lisas o con 1-3 papilas simples, 1 µm de alto (sur de Perú) … T. arequipensis

2. Células superiores y mediales de la lámina con (2)4-6(10) papilas simples o bifurcadas, 2.5-5 µm de alto … 3.

3. Células de la superficie ventral de la costa redondeadas a rectangulares, (15)22.5-25(32) µm de longitud, abultadas, dispuestas en 1-2 capas (en corte transversal) … T. atrovirens

3. Células de la superficie ventral de la costa cuadradas, rectangulares, algunas veces subcuadradas, raras veces oblatas o redondeadas (7.5)10-17,5 µm de longitud, no abultadas, dispuestas en 1 capa (en corte transversal) … 4.

3. Células mediales de la lámina 12.5-20 µm de ancho; dientes del peristoma aprox. 500 µm de largo, espiralmente torcidos (Peru?) … T. vahliana

3. Células mediales de la lámina (5)7.5-12.5(15) µm de ancho ; dientes del peristoma aprox. 100-132 µm de largo (registrado en Perú) … T. platyphylla