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Published In: Bryologia Europaea 1: 164 (fasc. 41. Monogr. 4). 1847. (Bryol. Eur.) Name publication detail
 

Project Name Data (Last Modified On 4/12/2011)
Acceptance : Accepted
Project data     (Last Modified On 4/12/2011)
Discussion:

Campylopus fragilis has enlarged hyaline cells throughout the leaf base, undifferentiated alar cells, and at times brood leaves clustered at the stem apices. The brood leaves in C. fragilis are identical to those found in Brothera leana (Sull.) C. Müll. Indeed, Bartram (1949) and Frahm (1991a) reported B. leana from Central America on the basis of small collections of C. fragilis that consist almost entirely of brood leaves. In B. leana the costa has a ventral and dorsal layer of hyalocysts, a median layer of stereid cells, and the upper leaf cells are long rectangular. Campylopus asperifolius has brood leaves borne on stalks, larger leaves, and firm-walled basal cells.

Campylopus tallulensis differs in having the costa in cross section with stereid bands extending to the margin, a more well-developed area of hyaline basal cells that continue up the margins of the leaves forming a V-shaped pattern, and better, although still weakly, developed alar cells. In C. fragilis the costal stereid bands do not reach the margins of the costa, which consists of 2–3 layers of enlarged, thin-walled, hyaline cells, the inner and outer hyaline basal cells end at about the same level in the leaf, and the alar cells are undifferentiated.

Illustrations: Grout (1937, Pl. 48 B); Bartram (1949, Fig. 19 E–F); Frahm (1978, Pl. 10); Smith (1978, Fig. 74 1–4); Frahm (1991a, Fig. 64). Habitat.
Habitat: On humus, rotting logs and soil; 2500–3130 m.
Distribution in Central America: GUATEMALA. Alta Verapaz: Croat 41757 (MO); Chimaltenango: Standley 58745a (FH); Huehuetenango: Sharp 4939a (FH); Quezaltenango: Sharp 1995 (FH); Quiché: Sharp 2484 (MO, NY, US); San Marcos: Croat 40918 (MO); Totonicapán: Standley 84107 (F). HONDURAS. Comayagua: Allen 12310 (MO, TEFH); Lempira: Allen 11238 (MO, TEFH); Morazán: Standley & Williams 428 (F); Olancho: Allen 12765 (MO, TEFH); Yoro: Allen 13557 (MO, TEFH). COSTA RICA. Alajuela: Brenes 17113 (F); Guanacaste: Brenes s.n. (FH); San José: Crosby & Crosby 6128 (MO).
World Range: Western Canada, Southeastern U.S.A.; Mexico; Central America; Caribbean, Western and Northern South America; Northern, Southwestern, Middle, and Southeastern Europe; Soviet Far East, China, Eastern Asia,; Indian Subcontinent; Macaronesia, Northeast Tropical, West-Central Tropical, East Tropical, South Tropical, and Southern Africa.

 

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Campylopus fragilis (Brid.) B.S.G., Bryol. Eur. 1: 164. 1847. Dicranum fragile Brid., J. Bot. 1800(2): 296. 1801. Type. Germany, Erlangen, Hoffmann s.n.

Plants small, tufted, green to yellow-green; stems usually densely tomentose, 0.6–3.5 cm long. Leaves 3–5 mm long, erect-patent, more or less flexuous above when dry, lanceolate, oblong below, subulate above, serrulate toward the apex; costa ending at the apex, in cross section 2/3 the leaf width, lightly ribbed at back, guide cells well developed, dorsal stereids present, ventral hyalocysts present and larger than the guide cells, at base the dorsal stereid band not as wide as the costa, the margins of the costa therefore of 2–3 layers of enlarged hyaline cells; upper cells rhombodial; basal cells rectangular, hyaline, thin-walled, bulging, in dried material frequently remaining full of air on wetting, marginal basal cells narrower but not forming a distinct border, cells just above the hyaline basal cells firm-walled, quadrate; alar cells not differentiated. Brood bodies minute, slender, at times clustered at the stem tips. Setae 5–8 mm long, reddish yellow; capsules erect to slightly curved when dry, 1.0–1.2 mm long; opercula shortly rostrate, 0.5 mm long. Calyptrae 1.0–1.2 mm long, fringed at base.

 

 

 
 
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