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Published In: Genera Plantarum 235–236. 1789. (4 Aug 1789) (Gen. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: S.M.H. JAFRI & M. QAISER
Contributor Institution: Herbarium, Department of Botany, University of Karachi, Karachi
General/Distribution: A family of 26 genera and nearly 200 species, distributed mostly in temperate and subtropical regions of northern hemisphere, including western north America and eastern Asia. Only 7 genera and 22 species have been reported from West Pakistan.
Comment/Acknowledgements: Acknowledgements: We are grateful to the United States Department of Agriculture for financing this research under P.L. 480. Thanks are also due to Messrs. B. L. Burtt and J. Cullen of Edinburgh, for their helpful suggestions.


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Annual, biennial or perennial herbs, very rarely shrubs or small trees, usually with milky or coloured latex. Leaves usually alternate, rarely opposite or whorled, exstipulate, mostly dissected, pinnately or palmately lobed. Infloresence basically cymose, mostly solitary, rarely subumbellate or paniculate. Flowers usually showy and large, bisexual, regular, hypogynous. Calyx of 2-3 free sepals, rarely completely united, imbricate, caducous. Petals 4-6 (rarely 8-12, or absent), in 1-2(-3) whorls, free, imbricate, sometimes crumpled in bud. Nectaries absent. Stamens indefinite in several whorls (only 4 stamens in subfam. Hypecoideae) free; anthers dithecous, dehiscing lengthwise; filaments filiform to winged and often coloured. Carpels 2-more, fused; ovary unilocular, rarely bilocular, usually with 2-several, incomplete locules; ovules numerous on parietal placenta (rarely solitary and basal, as in Bocconia ), anatropous or campylotropous; style 1 or absent; stigmas as many as the carpels, opposite or alternate with placenta. Fruit mostly a capsule, dehiscing by pores or valves, rarely indehiscent; seeds many, small, with a crested or smooth raphe or arillate; embryo minute; endosperm oily and mealy.

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1 Stamens numerous; inner petals not tripartite.(1. subfam. Papaveroideae) (2)
+ Stamens 4; inner petals ± tripartite (2. subfam. Hypecoideae) Hypecoum
2 (1) Fruit linear, at least 10 times as long as broad (3)
+ Fruit oblong or semiglobose, less than 10 times as long as broad (5)
3 (2) Latex yellow. Calyx not forming a hood. Receptacle not prominently hollowed (4)
+ Latex watery. Calyx forming a hood (which is detached and pushed all over the bud as the petals expand). Receptacle prominently hollowed Eschscholtzia
4 (3) Stigmas 2 conical. Fruit with 2 horns at apex, 2 valved Glaucium
+ Stigmas 3-4, capitate. Fruit without horns, 3-4 valved Roemeria
5 (2) Style present or inconspicuous. Stigma not discoid. Capsule opening by usually short valves (6)
+ Style absent. Stigmas discoid. Capsule opening by pores just beneath the persistent stigmas Papaver
6 (5) Style distinct, conspicuously broad and large. Stigma forming a globular mass over the ovary Meconopsis
+ Style inconspicuous (rarely very short). Stigma not forming a globular mass over the ovary Argemone
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