Lateral nectar glands usually in pairs, rarely annular, each semi-lunar or fragmentary; middle present or absent. Hairs simple, branched, glandular or absent. Stamen filaments linear, not appendaged. Ovary sessile, 1-4-ovuled, often subglobose; stigma capitate or shortly bilobed, rarely with erect or spreading lobes. Fruits indehiscent, neither beaked nor segmented; valves usually hard, often appendaged. Radicle incumbent or accumbent.