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Published In: Dictionnaire des Sciences Naturelles, ed. 2, 25: 321. 1822. (Dict. Sci. Nat. (ed. 2)) Name publication detailView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 5/13/2022)
Acceptance : Accepted
Project Data     (Last Modified On 9/3/2020)
Contributor Text: Abdul Ghafoor
Contributor Institution: Centre for Plant Conservation, University of Karachi – Pakistan
Synonym Text:

Lomatolepis Cass. in Cuvier, Dict Sci. Nat. 48: 422. 1827;  Rhabdotheca Cass. in Cuvier, Dict. Sci. Nat. 48: 424. 1827. Microrhynchus Less., Syn. Gen. Comp.: 139. 1832; Brachyrhamphus DC., Prodr. 7: 176. 1838; Zollikoferia DC., Prodr. 7: 183. 1838 nom. illeg. [non Zollikoferia Nees in Bluff & Fingerhuth, Comp. Fl. German. 2:305. 1825]; Heterochaena Fresen. in Mus. Senkenberg. 3: 74. 1839; Sonchus subg. Acanthosonchus Sch.Bip. in Webb & Berthelot, Hist. Nat. Iles Canaries 3 (2,2): 427. 1849; Paramicrorhynchus Kirp. in Komar., Fl. URSS 29: 725. 1964; Pavlov, Fl. Kazakh. 9: 510. 1966; Hexinia H. L. Yang in Liu, Fl. Desert. Reipubl. Pop. Sin. 3: 472. 1992.


 

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Annual to perennial herbs or rosetted shrublets or spinescent subshrubs with tap roots or rootstocks. Stem  weak, procumbent to ascending or erect and densely divaricately branched with stiff  shoots. Leaves  usually in basal rosette in young plants, sessile, entire to pinnatifid, cauline leaves amplexicaul, auriculate, mostly sinuate-dentate to pinnately dissected, rarely bipinnatifid, with white cartilaginous teeth. Capitula (5-) 7 - 40 (-100)-flowered, mostly numerous, borne in corymb-shaped or panicle-like synflorescence, very rarely capitula solitary;  peduncles few to several bracteate, usually patent-erect, becoming indurate and  ± spinescent after fruit fall, rarely very small. Involucre cylindrical, rarely ± globose. Phyllaries multiseriate, ± imbricate, much shorter, outer ½ to ¾ as long as inner subequal, linear-lanceolate to ovate, acute. Receptacle naked, pitted, the pits often toothed or ciliate margined. Florets with apically hairy corolla tube and ligules usually pale-yellow to yellow, narrowly spathulate to linear. Anther tubes obtuse above and with sagittate appendages at the base. Style branches long, slender, yellow with yellow or blackish hairs. Cypselas ± cylindrical to angular-prismatic, truncate or rarely with very short beak at the top, mono- or dimorphic, 4-10-ribbed, outermost cypselas often ± compressed and curved, glabrous or papillate, with smooth ribs or various types of transverse wrinkles. Pappus mono- or dimorphic, outer soft, cottony hairy, inner setaceous  hairy or all same, deciduous or persistent.

A genus with 54 species (Mabberley, 2008), distributed  in South Europe, Africa, Central, South and S W Asia and China. Represented in Pakistan by 11 species.

 

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1.

+

All cypselas with 5 primary ribs and 10 seconday ribs, basally 5-notched and with 5 projections.                            

 

 2

 

–             

Innermost cypselas with 4 primary and 8 secondary ribs, basally 4-notched with 4 projections. Peripheral cypselas  with 5 primary and 10 secondary ribs, basally 5-notched with 5 projections.           

               

 

 

 

 3

2.

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Synflorescence indurate to lignified, divaricately and intricately branched. Peduncles transformed  to spines or thorns after shedding of capitula and  buds. Young shoots and involucre beset with whitish mealy tomentum or glabrous. Marginal cypselas densely papillate  .                                              

 

 

 

 

 

 1. L. acanthodes

 

 

Synflorescence not indurated or lignified. Peduncles never transformed to spines or thorns after fall of capitula and buds. Marginal cypselas never densely papillate.              

 

 

 

 

2. L. aspleniifolia

3.

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Peripheral cypselas always papillate.   Pappus either persistent or deciduous   without disc.        

 

 

8

 

 

Peripheral cypselas always glabrous and mostly wrinkled. Pappus either persistent or  deciduous with disc.   

 

 

4

 

4.

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Pappus homomorphic, with setaceous rays only, deciduous with the disc.

 

 

                5

 

 

Pappus dimorphic, persistent, rays downy and setaceous.

               

 

6

 

5.

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Annual herbs with ± globose and subsessile 25-50-flowered capitula grouped in fascicles of (2-) 5-8 in a monopodial cyme or in centre of leaf-rosette in acaulescent plants. Anther    tube excl. appendages less than 1.5 mm long. Cypselas broadly winged. Fruiting receptacle c. 4 mm across.   

 

 

 

 

 

 

 7. L. capitata

 

 

Perennial herbs with cylindrical, pedunculate 16-27-flowered capitula grouped in fascicles borne in a dichasial panicle-like cyme. Anther  tube excl. appendages 2 – 3 mm long. Cypselas ribbed, wingless. Fruiting receptacle 1.5 – 3 mm across.   

 

 

 

 

 

 6. L. procumbens

 

6.

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Broom-like subshrub with indurated,   fastigiated leafless,long-articulate, virgate flowering axes.

 

 

4. L. oligocephala

 

Annual or perennial herbs, not broom-like

 but virgate or divaricately branched synflo-rescence and leafy or leafless flowering axes.

 

 

 

7

 

7.

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Perennial herbs with basal leaf rosette and

spike-like distinctly second synflorescence. Capitula 8-14-flowered. Ligules 7 – 8 mm long.  Anther tube excl. appendages 2.5 mm long.  Peripheral cypselas truncate, beakless, pale   to brownish. Pappus 4 – 5 mm long.      

 

 

 

 

 

 

 3. L. secunda

 

 

Annual herbs, with basal leaf rosette and more or less leafless shoots or foliaceous all over. Capitula 18-35-flowered. Ligules 5.5 – 6 mm  long.Anther tube ± 1.5 mm long. Cypselas shortly beaked, greyish to black. Pappus 7 – 8 mm long.

 

 

 

 

 5. L. intybacea

 

8.

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Capitula 25-30-flowered. Peripheral cypselas with a distinct glabrous short beak or cuspid and densely  papillate body.  Receptacle at fruiting stage c. 2 mm across.

 

 

 

8. L. quettaënsis

 

 

Capitula usually more than 30-flowered. All cypselas without a beak or cuspid. Receptacle at fruiting stage more  than 2.5 mm across.               

 

 

 

9

 

9.

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Style branches with blackish sweeping hairs. Inner ripe cypselas with distinct secondary ribs and papillate in transverse rows.

 

 

 

 9. L. stenocephala

 

 

Style branches with yellow sweeping hairs. Inner ripe cypselas without seconday ribs and smooth to weakly wrinkled in transverse rows.

 

 

 

10

 

10.

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Plants glaucous-green, mostly less than 20 (-25) cm tall. Leaves shallowly pinnatifid, with pinnatifid auricles at the base. Ligules 10 – 14 mm long.

 

 

 

 11. L. platyphylla

 

 

Plants glaucous-greyish, mostly more than 20 cm tall. Leaves deeply uni- or bipinnatifid, with cartilaginous teeth on sinuate margins. Ligules 7 – 10  mm long.

 

 

 

 

10. L. mucronata

 

 
 
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