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Published In: Annales des Sciences Naturelles (Paris) 2: 346. 1824. (Jul 1824) (Ann. Sci. Nat. (Paris)) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: SULTANUL ABEDIN and S.I. ALI
Contributor Institution: Department of Botany, University of Karachi, Karachi.
General/Distribution: A family of 16 genera and about 500 species distributed in tropical regions of both hemispheres. Only 3 genera and 4 species are known from W. Pakistan.
Comment/Acknowledgements: Acknowledgements: We are grateful to the United States Department of Agriculture for financing this research under P.L. 480. Thanks are also due to Mr. B.L. Burtt (Edinburgh), Mr. E. Milne-Redhead (Kew) and Dr. S.A. Faruqi (Karachi) for their helpful suggestions.


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Shrubs or trees with resin or aromatic oil. Leaves alternate, rarely opposite or fascicled, usually pinnately compound or decompound, l-3 (—or more) foliolate, imparipinnate. Flowers in panicles, solitary or sometimes fascicled, bisexual or unisexual (plants then diclino or polygamodioecious), actinomorphic and hypogynous. Calyx 3-6 fid or partite forming a tube, lobes imbricate or valvate. Petals 3-6, free, rarely connate below, imbricate or valvate. Disc annular or cupular, rarely absent, free or adnate to the calyx tube. Stamens in l-2 whorls, as many as or twice the number of petals, free, or rarely connate at the base; anthers usually versatile, dithecous, with longitudinal dehiscence. Carpels 2-5, syncarpous; ovary sessile or stipitate, 2-5-loculed; each locule 2(-1) ovuled; placentation axile; style 1 or absent; stigma 2-5-lobed or undivided. Fruit drupaceous, indehiscent with 2-5 pyrenes or dehiscent. Seeds usually with membranous testa and without endosperm.

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1 Receptacle cup or bell shaped; sepals valvate. (2)
+ Receptacle flat; sepals imbricate. Bursera
2 (1) Sepals 5, not persistent; petals 5; stamens 10; stigma 4-5 lobed; fruit with stones never growing together; disc lining the tube of the calyx. Garuga
+ Sepals 4, persistent; petals 4; stamens 8; stigma 2 lobed; fruit with stones growing together; disc not lining the tube of the calyx. Commiphora
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