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Published In: Journal für die Botanik 1800(2): 299. 1801. (J. Bot. (Schrader)) Name publication detail
 

Project Name Data (Last Modified On 3/13/2009)
Acceptance : Accepted
 

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29. Syntrichia Brid.   赤藓属 chi xian shu
J. Bot. (Schrader) 1(2): 299. 1801.
 
Plants usually medium-sized to large, sometimes rather robust, green to reddish brown, in loose or dense tufts. Stems erect, simple or irregularly branched, radiculose at base; central strand usually present, rarely absent. Leaves appressed or contorted-curved to somewhat crisped when dry, spreading or sometimes reflexed when moist, oblong-ligulate to spathulate, or narrowly ovate-lanceolate, rounded-obtuse to broadly acute or mucronate at apex, upper lamina broadly channeled or keeled, weakly sheathing at base; margins often recurved, crenulate by projecting papillae, usually forming a differentiated border, consisting of several rows of thick-walled, or sometimes elongated cells, yellowish or brownish, border rarely absent; costa rather stout, often reddish brown, excurrent, ending in a short to rather long awn, often denticulate on the back or at tips, costa rarely percurrent or ending below the apex, in cross section 1 dorsal stereid band present (which is exposed, not covered by parenchymatous epidermal cells) or sometimes the ventral stereid band very weakly developed; upper leaf cells subquadrate to rounded-hexagonal, each cell densely covered with several C-shaped papillae; basal cells distinctly differentiated, rectangular, thin-walled, hyaline, smooth. Autoicous or dioicous. Setae elongate, straight, yellowish brown; capsules erect or slightly inclined, cylindrical; peristome teeth 32, linear or filiform, twisted, papillose, rarely erect; basal membrane high; annuli differentiated; opercula conic-rostrate, with a long, oblique beak. Calyptrae cucullate, smooth. Spores spherical, small, yellowish green or reddish brown, smooth or finely papillose.
 
 

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We follow Zander’s (1993) system, separating Syntrichia from Tortula. Six species of Syntrichia are recognized in this study.
 
1. Leaves rounded-obtuse, mucronate; costa percurrent or somewhat excurrent.............................. 2. S. fragilis
1. Leaves acute or broadly acute; costa rather long-excurrent, ending in an awn.................................................. 2
2. Leaf margins usually rather plane or only slightly recurved; costa smooth on the dorsal surface, ending in a smooth or distantly denticulate awn......................................................................................................... 3
2. Leaf margins rather strongly recurved to revolute; costa dentate on the dorsal surface, ending in a coarsely serrate awn.......................................................................................................................................... 4
3. Awns hyaline, distantly denticulate; upper leaf cells pentagonal or hexagonal, rather thin-walled; basal membrane low, less than 1¤4 peristome teeth length.......................................................... 3. S. longimucronata
3. Awns reddish brown, smooth; upper leaf cells rounded-hexagonal, rather thick-walled; basal membrane high,
ca. 1¤3 the peristome teeth length...................................................................................................... 6. S. sinensis
4. Plants usually less than 1.5(–2) cm high; upper lamina often bistratose in cross section...........1. S. caninervis
4. Plants usually larger than (2–)3(–8) cm high; upper lamina unistratose in cross section........................ 5
5. Autoicous (synoicous); basal leaf cells lax, thin-walled, short-rectangular to quadrate at the margins, ca. twice as long as wide; stem with a central strand........................................................................... 4. S. princeps
5. Dioicous; basal leaf cells firm, somewhat thick-walled, narrowly rectangular to quadrate of narrowly rectangular at the margins, at least three times as long as wide; stem without a central strand.... 5. S. ruralis
 
 
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