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Published In: Species Plantarum 1: 477. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 5/12/2022)
Acceptance : Accepted
Project Data     (Last Modified On 5/25/2018)
Contributor Text: H. McAllister
Contributor Institution: School of Biological Sciences, University of Liverpool, Liverpool. U.K.
Comment/Acknowledgements:

SORBUS AFF. CASHMIRIANA SP. NOV.

A species from Pakistan, North West Frontier Province, Swat Division, Gabrak Tal, 2600 m. 1983, Berger 5712, is a tetraploid (2n=68) apimictic microspecies very similar to S. cashmiriana and probably not distinguishable on a herbarium sheet. However, its flowers are almost white and conspicuously less pink than in S. cashmiriana, and its leaflets are slightly smaller.

Herbarium specimen in LIV. Cultivated specimens in Berlin, Dahlem; University of Liverpool Botanic Gardens, Ness; Reykjavik City Botanic Garden.

S. aff. cashmiriana

Chitral-Drosh, 35°21’17.1”, 71°48’37.7”, Lowari top, 3276 m, 5.7.2006, rocky slope, very rare, Haidar Ali 4479 (KUH).

Buds rufous-hairy. Leaves up to c. 12 cm, leaflets narrow, oblong–elliptic (cf. ovate-elliptic in S. cashmiriana), 9-10 pairs, closely inserted (c. 1cm between nodes on rachis, (cf. c. 1.5 cm in S. cashmiriana), leaflets coarsely toothed. As fruits are immature, ultimate fruit size is uncertain and, as they are more densely clustered than would be expected for S. cashmiriana, it may be quite unrelated distinct taxon. Fruits have not been dissected.


 

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Tall (to 30 m) to small trees or shrubs; bark smooth or flaky; twigs very variable, from very stout and stiff to thin and flexible. Buds, shortly ovate to conic, with a variable number of bud scales visible. Leaves simple or pinnately divided, spirally arranged. Inflorescence a panicle, corymbose to pyramidal; flowers more or less perigynous, calyx lobes triangular deltoid, sometimes gland toothed but otherwise entire, green with leaf-like texture (with distinct midrib) or fleshy (with midrib not distinct) in fruit; petals white to pink to crimson, clawed; stamens usually 20; carpels 2—5, remarkably variable in degree of fusion with receptacle and to one another. Fruit a berry-like pome, green to orange-red or crimson to white, from 5.5-15 mm; seeds usually flattened ovoid.

In Pakistan subgenus Aria is represented by S. lanata, while subgenus Sorbus is represented by S. tianshanica of the red-fruited group and five apomictic microspecies of three of the species groups of the white-crimson fruited species group. The three named and one un-named microspecies of the S. cashmiriana group may be rather difficult to distinguish in the herbarium, though the living trees are not difficult to identify. More microspecies, especially of the S. cashmiriana group, are likely to be present.

 

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1.

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Leaves simple but pinnately lobed.

1.  S. lanata

 

Leaves pinnately divided with 5-12 pairs of distinct leaflets and one terminal leaflet.

 

2

2.

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Fruit cherry-pinkish-red, flesh yellow when fresh with obvious chromoplasts (x 400), carpel apices free, white-hairy and unfused within calyx; bark on young twigs very glossy; leaves glabrescent, leaflets narowly elliptic 

                   

 

2. S. tianshanica

 

Fruit white to pink variably flushed with crimson, flesh white when fresh with no plastids visible (x 400), carpel apices densely (S. cinereopubescens) or slightly white hairy, more or less fused except in S. cinereopubescens

 

3

3.

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Leaflets glaucous and minutely papillose beneath (x 70), oblong to oblong-ovate, blunt or apiculate. Fruit firm, to 11mm across, often broader than long, calyx lobes in depression at fruit apex (i.e. fruit apple-shaped) Flowers white, style about 2.5mm.                                                                                 

 3. S. foliolosa

 

Leaflets non-papillose beneath, ovate to lanceolate, more or less acute; styles longer.

 

4

4.

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Twigs thick, stiff; leaves persistenly hairy beneath on rachis and leaflet midribs; fruit often broader than long to 13mm wide, styles about 3 mm. Carpel apices hairy, distantly inserted on densely white-hairy carpel apices.

                       

 

4.  S. cinereop- ubescens

 

Leaves glabrescent; leaflets ellipsoid-lanceolate, acute; fruit very soft, large, often more than 12mm across, with fleshy calyx lobes protruding somewhat at fruit apex giving a fruit which is longer than broad with an inverted pear shape; carpel apices slightly hairy; styles usually longer than 3.5mm (S. cashmiriana agg.).

 

5

5.

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Fruit pink                                                                                                             

5. S. rosea

 

Fruit white.

6

6.

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Carpels completely fused at apices; autumn colouration yellow.

6. S. cashmiriana

Grooves visible between carpels at apices; autumn colouration yellow and orange-red.                                                                                        

 

7.  S. gilgitana

 
 
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