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Published In: Synopsis Universalis Analytica Generum Plantarum 2: 395. 1794. (Syn. Univ. Analytica Gen. Pl.) Name publication detail

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: YASIN J. NASIR
Contributor Institution: National Herbarium, Pakistan Agricultural Research Council, Islamabad.
General/Distribution: 24 genera and c.1000 species, mainly in the N. Hemisphere. Represented in Pakistan and adjoining areas by 8 genera and 60 species. Various cultivars of the genus Cyclamen L., are occasionally grown as ornamental plants.
Comment/Acknowledgements: Acknowledgements: I am highly indebted to the Directors/Curators of the following herbaria for the loan of herbarium specimens: B, BG, BM, BR, CGE, DD, E, G, GH, GL, H, ISL, K, KUH, L, LAH, LD, LE, LINN, LIV, LWG, M, MICH, 0, P, PES, PPFI-B, PPFI-M, PAN, RAW, S, UPS, US, W, WRSL, WU, Z.

I am thankful to Mr. Airy Shaw Smith (Kew) and Mr. I.C. Hedge (Edinburgh) for the latin diagnoses to the new taxa. For information concerning the typification of Wallich’s Types I am grateful to Mr. A. Radcliff Smith (Kew) and to Miss Sylvia Gould (BM). I would also like to thank the following individuals for their help and cooperation in various ways: Dr. F.R. Fosberg, Dr. D. Nicolson, Dr. J. Nowicke (Smithsonian Institute, Washington), Mr. C.C. Townsend and Dr. T.A. Cope (KEW), Mrs. J. Woods (E), Mr. A Chater (BM), Dr. M. Qaiser, Mr. M. Shariq and Dr. Usman Ghani Khan (Karachi University), Dr. M. Afzal (Pakistan Science Foundation, Islamabad) and Dr. Shahina Ghazanfar (formerly at PARC, Islamabad).

The financial assistance received from the United States Department of Agriculture, under PL-480 is gratefully acknowledged.


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Perennial or annual terrestrial herbs, sometimes suffruticose, often with underground parts as rhizomes or tubers. Stem erect or prostrate, simple or branched, sometimes as stolon or runners or assuming a cushion like habit. Leaves simple, radical, in basal rosettes or cauline, alternate, opposite or verticellate, exstipulate, entire or variously lobed, sessile to petiolate, sometimes farinose, revolute, involute or conduplicate in bud. Flowers bisexual, regular, 5(-7-9)-merous, axillary solitary or few or disposed in panicles, racemes or spikes, sometimes in verticels, umbellate or in few flowered or dense capitate heads, fairly large and showy or small and inconspicuous; scape well developed or obsolete. Bracts present, sometimes leafy, often prolonged downwards to form a spur. Bracteoles absent. Pedicels variable in length and may elongate after antheses. Calyx 1/3rd to 2/3rd cleft, persistent. Corolla campanulate, rotate or infundibuliform, 5-lobed, imbricate or convolute, sometimes absent or nodding; tube long or short. Throat may be constricted and sometimes obstructed with hairs. Stamens 5, epipetalous and opposite the corolla lobes. Slender or scale-like staminodes, alternating with corolla lobes present in Samolus. The anther dehiscence introrse and by longitudinal slits; filaments free or adnate at the base of the corolla. Carpels 5, syncarpous. Ovary superior or semi-inferior (Samolus), with one to many ovules on basal or free central placentae; ovules semi-erect or erect (Samolus), usually bitegmic. Style and stigma simple, the latter capitate. Fruit a capsule dehiscing by 5(-10) valves which are opposite the sepals, or irregularly or by a lid (Anagallis). Seeds few to many, testa smooth to vesiculose or papillose, often reticulate or faintly so.

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1 Corolla well developed. Calyx not petaloid (2)
+ Corolla absent. Calyx petaloid 6 Glaux
2 (1) Ovary superior, ovules semi-erect; the inner integument 3 cell thick. Seeds .02-2.7 mg (3)
+ Ovary semi-inferior, ovules erect; the inner integument 2 cell thick. Seeds less than .05 mg 8 Samolus
3 (2) Plants usually scapigerous with leaves basal or in rosettes. Corolla infundibuliform to salver-shaped, tube usually long (short in Androsace), aestivation imbricate (4)
+ Plants escapose, with leafy stems. Corolla rotate, tube short, aestivation convolute (7)
4 (3) Herbaceous plants (sometimes with a ± woody base in some Androsace sp.). Corolla tube not curved. Farina, if present, of a mealy nature (5)
+ Suffrutescent herbs or semi-herbs. Corolla tube often curved. Farina if present, usually of the woolly type 2 Dionysia
5 (4) Flowers not nodding (except in some Primula species). Anthers not apiculate. Pollen grains with a microreticulate to reticulate tectum (6)
+ Flowers nodding. Anthers apiculate. Pollen grains with a granular tectum 4 Cortusa
6 (5) Plants pulvinous or of a loose caespitose habit. Flowers monomorphic. Corolla tube short, about equalling the calyx, urceolate; throat constricted. Pollen grains with a distinct pore. Capsule globose, included in calyx. Seeds up to 50 per capsule, usually larger (0.8-3.0 mm long) 3 Androsace
+ Plants rarely pulvinous. Flowers often dimorphic. Corolla tube elongated and usually exceeding the calyx; throat not constricted. Pollen grains with an indistinct pore. Capsule globose or cylindrical, usually exserted. Seeds (up to 150 per capsule, usually smaller (0.3-2.5 mm long) 1 Primula
7 (3) Capsule dehiscence valvular 5 Lysimachia
+ Capsule dehiscence operculate 7 Anagallis
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