Lateral nectar glands various, often annular or semi-annular, simple or variously lobed; middle glands present or absent. Hairs simple, branched or absent. Sepals subequal, erect or subspreading. Stamen filaments without append-ages, rarely toothed. Ovary sessile or with short gynophore; stigma capitate, often slightly bilobed. Siliquae usually linear (rarely elliptic or sub-globose sillicula-like), usually compressed or flattened parallel to the septum, dehiscent (rarely tardily or hardly dehiscent, broadly septate (rarely septum incomplete or obsolete); seeds 1-2-seriate, generally not muciloginous when wet.; radicle usually accumbent.