Home Flora of Pakistan
Home
Name Search
Families
Genera
Species
District Map
Grid Map
Inventory Project
Nepeta L. Search in The Plant ListSearch in IPNISearch in Australian Plant Name IndexSearch in Index Nominum Genericorum (ING)Search in NYBG Virtual HerbariumSearch in JSTOR Plant ScienceSearch in SEINetSearch in African Plants Database at Geneva Botanical GardenAfrican Plants, Senckenberg Photo GallerySearch in Flora do Brasil 2020Search in Reflora - Virtual HerbariumSearch in Living Collections Decrease font Increase font Restore font
 

Published In: Species Plantarum 2: 570. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: I.C. Hedge
General/Distribution: Over 200 species mainly in Eurasia. The largest labiate genus by far in Pakistan, with a mixture of some very clear-cut species and others extremely polymorphic. Many of the difficulties of classification stem from the paucity of reliable characters. To mention only two, indumentum and calyx teeth can vary greatly within an individual species. One of the few characters that seems reliable is nutlet size, shape, surface texture and areole length. Male-sterile flowers occur in a number of species; hybridization and introgression probably also play a role in the complexities of classification. Some species have far from adequate types. Many outstanding problems still remain among the Pakistan species and an improvement on the existing classification will have to be preceded by detailed studies in the field; and collation with the taxonomic situation in adjacent areas. Kashmir is an area where field studies are particularly needed; about half of the species in this account grow there.
Comment/Acknowledgements: It is almost impossible to formulate a truly satisfactory and practical key to the species; inevitably some of the contrasts in the couplets are not clearcut.

Species of doubtful status or doubtfully in our area

Nepeta ciliaris Wall. ex Benth. in Wall. [Cat. 2082. 1829], Pl. As. Rar. 1: 64. 1830; Stewart, Ann. Cat. Vasc. Pl. W. Pak. & Kashm. 619. 1972.

This is a problematical species nomenclaturally and taxonomically. The specimen of Wallich Cat. No. 2082 (collected by Royle) at Kew seems not to agree with the original description of Nepeta ciliaris and in my opinion is Nepeta distans. The name Nepeta cilians has generally been applied to plants related to Nepeta leucophylla Benth. and also to Nepeta hindostana, but I have not seen any specimens of it from Pakistan.

Nepeta glomerulosa Boiss., Diagn. Pl. Or. Nov. Set. 1, 5: 21. 1844; Mukerjee in Rec. Sot. Surv. Ind. 14, 1: 130. 1940; Stewart, Ann. Cat. Vasc. Pl. W. Pak. & Kashm. 621. 1972.

Plants from Pakistan so-named are better identified as Nepeta juncea (q.v.), but they are part of a difficult complex stretching west to Iran. Nepeta glomerulosa is the earliest specific epithet.

Nepeta leucophylla Benth., Lab. Gen. et Sp. 476. 1834; Kitamura, Pl. W. Pak. & Afghan. 128. 1964; Stewart, Ann. Cat. Vasc. Pl. W. Pak. & Kashm. 623. 1972.

I have seen no Pakistan specimens which match this more eastern mainly Himalayan species. It is a close ally of Nepeta ciliaris.

Nepeta micrantha Bunge in Ledeb., Fl. Alt. 2: 401. 1830; Hedge & Lamond in Notes Roy. Bot. Gard. Edinb. 28: 118. 1968; Stewart, Ann. Cat. Vasc. Pl. W. Pak. & Kashm. 623. 1972.

The record of this annual in Pakistan seems to stem from Boissier (Fl. Orient. 4: 664. 1879) who cites Stocks 1023 from Baluchistan. The specimen of this number at Kew is Drepanocaryum sewerzowii (Regel) Pojark. Nepeta micrantha is mainly a C. Asiatic species.

Nepeta multicaulis S.K. Mukerjee in Journ. Ind. Bot. Soc. 19: 86, f. 6. 1940.

The description, drawing and type locality (Baltistan, Saltaro) suggest that this is a form of Dracocephatum stamineum , q.v., in which the stamens are not exserted. The type (Hunter-Weston, herb. Debra Dun 10266) has not been seen.

Nepeta spathulifera Benth. in DC., Prodr. 12: 380. 1848; Mukerjee in Rec. Bot. Surv. Ind. 14, 1: 126. 1940.

Mukerjee (l.c.) indicated this species from Baluchistan without giving details. Rechinger (Fl. Iran. 150: 200. 1982), for the external distribution of this Afghan-Pamir Alai annual, gave "Pakistan/Gilgit"; this possibly stems from his giving Nepeta gilesii, described from Gilgit, as a synonym of Nepeta spathulifera. In this account Nepeta gilesii is given as a synonym of Nepeta floccose.


 

Export To PDF Export To Word
Annual or perennial often aromatic herbs of varied habit and indumentum; hairs usually simple, rarely branched. Leaves simple, rarely pinnatisect, with entire, crenate or serrate margins, sessile or petiolate. Inflorescence of verticillasters, distant or crowded into spike-like or ovoid heads, or lax pedunculate cynics, borne in axils of upper leaves or not. Bracts shorter to longer than calyces. Flowers hermaphrodite or male-sterile. Calyx ± clearly 15-veined or ribbed, ± tubular, bilabiate, little changed in fruit, straight or curved; upper lip 3-dentate, lower 2-dentate, rarely both lips entire; tube glabrous within or pilose at throat; throat straight to strongly oblique. Corolla blue, violet, pink, yellow or white, bilabiate; tube straight or curved, included in or exserted from calyx; upper and lower lips relatively short. Stamens 4, didynamous, posterior pair longer than anterior; thecae usually diverging at an angle of 180°. Style unequally bibbed. Nutlets ellipsoid to obovoid, apically rounded, smooth or variously tuberculate, mucilaginous on wetting or not; areole (attachment scar) bibbed, of varied length.
 

Export To PDF Export To Word Export To SDD
Switch to bracketed key format
1.Annuals
2.Leaves bipinnatisect
Nepeta annua
2.Leaves undivided
3.Inflorescence of clearly separated verticillasters or of lax pedunculate cymes
4.Cymes pedunculate with clearly pedicellate flowers
6.Calyx c. 4.5 mm; nutlets clearly depressed-tuberculate; peduncles filiform
Nepeta graciliflora
6.Calyx 3.5-4 mm; nutlets inconspicuously depressed-tuberculate; peduncles not filiform
Nepeta hindostana
4.Verticiilasters borne in axis of leaves or bract-like upper leaves, ± sessile
7.Indumentum dense, of long white villous hairs; nutlets verrucose with a long U-V-shaped areole
Nepeta eriosphaera
7.Indumentum of short hairs; nutlets smooth, shiny, with a small basal areole
8.Leaves linear-elliptic; stems with some papillose glandular hairs
Nepeta daenensis
8.Leaves ovate to rhombic; stems with only eglandular retrorse hairs
Nepeta bracteata
3.Inflorescence of terminal ovoid heads
7.Indumentum dense, of long white villous hairs; nutlets verrucose with a long U-V-shaped areole
Nepeta eriosphaera
7.Indumentum of short hairs; nutlets smooth, shiny, with a small basal areole
8.Leaves linear-elliptic; stems with some papillose glandular hairs
Nepeta daenensis
8.Leaves ovate to rhombic; stems with only eglandular retrorse hairs
Nepeta bracteata
1.Perennials
9.All cauline leaves sessile, amplexicaul or not, or lowest stem leaves with a short, up to 3 mm petiole
10.Plants with a dense all-over indumentum of long and short capitate glandular hairs
11.Leaves deeply serrate-incised
Nepeta glutinosa
11.Leaves crenate
Nepeta adenophyta
10.Plants not as above, glabrous, eglandular pilose or vinous, or with glandular papillae
12.Verticillasters crowded in ails of uppermost, often lilac-blue, leaves
Nepeta coerulescens
12.Verticillasters forming elongated spikes not as above
13.Leaves pectinate-serrate, narrow or broad elliptic; indumentum of plant ± dense, villous, with or without glandular papillae
Nepeta elliptica
13.Leaves with entire or finely serrulate margins, linear to linear-elliptic; indumentum sparse or subabsent without glandular papillae
14.Calyx 11-16 mm; corolla 20-25 mm; pedicel up to 12 mm
Nepeta connata
14.Calyx 10 mm or less; corolla 12-15 mm; pedicel absent to 2 mm
15.Leaves up to 20 mm long; calyx 5-7 mm [Quetta]
Nepeta podostachys
15.Leaves 20-90 mm long; calyx 6-10 mm [NW Pakistan, Kashmir]
16.Leaves linear 1.5-5 mm wide; root- stock tuberous or fusiform
Nepeta linearis
16.Leaves linear-lanceolate to linear-elliptic, up to 10 mm wide; rootstock not as above
17.Spike longer and slender than that of N. nervosa, often interrupted near the base; leaves oblong-lanceolate, c. 25-50 x 12-20 mm
Nepeta campestris
17.Spike shorter than that of N. campestris, congested not interrupted near the base; leaves linear-lanceolate or linear-elliptic, up to 80 x 10 mm
Nepeta nervosa
9.At least the lower leaves clearly petiolate with a petiole more than 4 mm
18.Calyx with entire upper and lower rounded lobes
Nepeta suavis
18.Calyx with distinct teeth
19.Stems with an indumentum of distinct numerous stellate-dendroid hairs
20.Inflorescence of several ± distant verticillasters borne in axils of leaves or leaf-like bracts; calyx with a dense stellate-dendroid indumentum
Nepeta leucolaena
20.Inflorescence of mostly solitary terminal ovoid heads; calyx with pilose or villous simple hairs
Nepeta kokanica
19.Stems without or with few indistinct furcate or branched hairs (strong magnification needed)
21.Inflorescence of loose pedunculate cymes with clearly pedicellate flowers
22.Nutlets with a prominent U- or V-shaped areole extending c. 2/3 nutlet length
23.Leaves c. 7-10 x 7-10 mm, serrulate; peduncles up to 20 mm [NW Pakistan: Chitral]
Nepeta brachyantha
23.Leaves up to 25 x 25 mm, crenate-dentate; peduncles up to c. 10 mm [W Pakistan)
22.Areole of nutlet c. 1/3 or less the nutlet length
24.Calyx c. 4.5 mm; nutlets clearly depressed-tuberculate; peduncles filiform
Nepeta graciliflora
24.Calyx 3.5-4 mm; nutlets inconspicuously depressed-tuberculate; peduncles not filiform
Nepeta hindostana
21.Inflorescence of ± condensed pedunculate cymes, of distant verticillasters, ovoid heads or spikes
25.Corolla 20-25 mm long
26.Larger leaves 4-11 cm long
27.Corolla yellowish, c. 25 mm long
Nepeta govaniana
27.Corolla mauve to deep violet-blue, 20-25 mm long
Nepeta erecta
26.Larger leaves 15-3 cm long
28.Indumentum on inflorescence vinous, with flat eglandular multicellular hairs; stems ± uniformly leafy
Nepeta glechomifolia
28.Indumentum on inflorescence sparse, of short eglandular or glandular hairs; leaves mostly on lower part of stem
Nepeta pinetorum
25.Corolla 6-17 mm long
29.Verticillasters ± clearly separated from each other or uppermost loosely con-gested
30.Corolla c. 15 mm or more
31.Stems erect-ascending, slender in region of inflorescence; bracts subulate, 2-4 mm
Nepeta pinetorum
31.Stems erect, ± sturdy throughout; bracts linear 1/2 as long as to as long as calyces
Nepeta clarkei
30.Corolla less than 10 mm
32.Inflorescence paniculate
33.Basal leaves suborbicular-ovate, white tomentose to lanate; cymes mostly on horizontally spreading peduncles
Nepeta floccosa
33.Basal and lower leaves triangular-ovate with a greyish indumentum; peduncles not horizontally spreading
Nepeta cataria
32.Inflorescence unbranched or lower verti-cillasters shortly pedunculate, not panicu-late
34.Inflorescence of ± globose densely white-lanate-villose verticillasters; stems procumbent-ascending; nutlets with a U-V-shaped areole c. 3/4 length of nutlet
Nepeta lagopsis
34.Inflorescence not lanate-villose; stems erect; areole c. 1/3 length of nutlet
35.Bracts c. 1/3 length of calyx; calyx often purplish
Nepeta rugosa
35.Bracts 1/3 to as long as calyx; calyx pale green or straw-coloured
36.Leaves ovate-triangular, ± as long as broad; verticillasters up to c. 15 mm diameter in fruit
Nepeta praetervisa
36.Leaves oblong to oblong-ovate, longer than broad; verticillasters up to 10 mm in fruit
Nepeta juncea
29.Verticillasters forming ± continuous spikes or ovoid heads, sometimes lowermost verticillasters distant
37.Rootstock with a distinct tuber 8-20 mm in diameter [leaves green, broad triangular-ovate, 5-20 x 5-15 mm; calyx teeth subequal]
Nepeta raphanorhiza
37.Rootstock not tuberous
38.Scree plants with scale-like leaves on rootstock
39.Leaves cuneate, margins deeply incised with obtuse round lobes; at least some bracts as long as or longer than flowers
Nepeta longibracteata
39.Leaves cordate, margins crenulate; bracts ± equal to calyces
Nepeta kurramensis
38.Not scree plants; without scalelike leaves on rootstock
40.Leaves markedly discolorous or grayish white canescent
41.Leaves densely adpressed canescent with at least some branched hairs (strong magnification needed)
Nepeta kokanica
41.Leaves discolorous; all hairs unbranched
42.Nutlets smooth; outer bracts elliptic to obtrullate
43.Leaves c. 7-20 x 7-12 mm; corolla c. 12 mm
Nepeta discolor
43.Leaves up to 30 (-45) x 20 (-30) mm; corolla up to 17 mm
Nepeta eriostachys
42.Nutlets tuberculate; outer bracts ± linear
44.Calyx 9-10 mm; leaf apices obtuse
Nepeta schmidii
44.Calyx 6.5-8 mm; leaf apices ± acute
45.Perennial herb, 30-50 cm, leaves broadly ovate to triangular-ovate, 15-35 x 10-30 mm; inflorescence of numerous sessile or shortly pedunculate, many-flowered congested cymes forming an elongated or short spike interrupted below
Nepeta griffithii
45.Dwarf suffruticose herb, c. 10 cm high, leaves c. 15 x 18 mm, cordate-reniform; verticillasters condensed into a ± loose head or spike
Nepeta amicorum
40.Leaves ± green on both surfaces
46.Vertillasters crowded in axils of uppermost often lilac-blue leaves; lowermost leaves with a short petiole to 4 mm, all other leaves sessile
Nepeta coerulescens
46.Verticillasters not as above; lowermost leaves with petioles of 6 mm or more
47.Inflorescence a terminal ovoid head
48.Leaves serrate, acute
Nepeta paucifolia
48.leaves crenate, apically rounded or obtuse
49.Plants not aromatic, suffruticose, clump-forming, stems 15-30 cm; leaves ± broadly ovate, flabelliform or trullate, indumentum of simple or branched hairs; inflorescence of terminal congested ovoid heads, or sometimes with a distinct lower verticillaster
Nepeta kokanica
49.Plants aromatic, stem erect, c. 30 cm; leaves triangular-ovate, indumentum of simple hairs; inflorescence of 2-3 con-densed or separated verticillasters
Nepeta grata
47.Inflorescence an elongated oblong-cylindrical spike
50.Outer bracts at base of spikes ovate-lanceolate, often coloured
51.Leaves ± broad-ovate, 12-20 x 10-15 mm, with a rather dense short indumentum of retrorse hairs
Nepeta distans
51.Leaves triangular-ovate to elliptic, 30-60 x 20-30 mm, usually glabrous to minutely pilose
Nepeta laevigata
50.Outer bracts at base of spikes narrow lanceolate to linear
52.Leaf margin serrate to subpectinate
Nepeta subincisa
52.Leaf margin crenate to crenulate
53.Leaves linear-elliptic to elliptic, rarely with some leaves ternately pinnatisect; clusters of young leaves present in leaf axils
Nepeta paulsenii
53.Leaves oblong-ovate to oblong-triangular, simple; usually without clusters of young leaves in leaf axils
54.Leaves 10-20 x 8-15 mm, glabrous to pilose; inflorescence ± glabrous
Nepeta podostachys
54.Leaves 20-40 x 15-25 mm, villous; inflorescence ± white villous
Nepeta pubescens
5.Cauline leaves linear, upper spreading and recurved
5.Cauline leaves ovate to elliptic, upper spreading
Nepeta pungens
 

Lower Taxa
 
 
© 2024 Missouri Botanical Garden - 4344 Shaw Boulevard - Saint Louis, Missouri 63110