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Published In: Species Muscorum Frondosorum 178–187, pl. 42, f. 8–12; pl. 43–44. 1801. (Sp. Musc. Frond.) Name publication detailView in Botanicus

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Bryum is a large and extremely variable genus of acrocarpous mosses generally found on soil or humus. The plants often have a rosette growth form with the upper leaves larger and better developed than the lower leaves. The more robust species of Bryum, which often have a strongly rosulate habit, have recently been segregated into the genus Rosulabryum (Spence 1996). Typically the leaves are bordered by linear cells, have large rhomboidal to oblong‑rhomboidal upper cells, and percurrent to excurrent costae. Its capsules are usually pendent, more or less pyriform, and have well‑developed, often perfect, diplolepidous peristomes. Bryum is considered taxonomically difficult due to its usually small size and generally non‑descript, often technical characters. Adding to the difficulty, leaves taken from different parts of single plants may have dissimilar morphologies and character states. Within Bryum, groups of seemingly related forms can sometimes be recognized, but " .... a clear division into subgenera or sections is not indicated". (Andrews 1940). Characters that are important in the taxonomy of Bryum include: 1. leaf cell areolation (upper leaf cell length vs width; thickness of cell walls; basal leaf cell size and shape), 2. costal length; 3. leaf margin stance and border development, 4. plant sexuality, 5. leaf decurrency development, 6. capsule shape; 7. endostome development.


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Bryum Hedw. Sp. Musc. Frond. 178. 1801. 

Rosulabryum Spence, Bryologist 99: 222. 1996, syn. nov. 

Plants small to robust, in erect, dense or loose tufts, simple or forked. Leaves erect to erect‑spreading when dry, spreading when wet, ovate, ovate‑lanceolate, lanceolate, oblong-lanceolate, oval, elliptic, or suborbicular, often decurrent; apices obtuse, acute or acuminate, concave; margins plane, erect, or recurved, entire to serrate near apex; costa strong, subpercurrent to excurrent; cells smooth, thin‑ or thick‑walled, quadrate to long‑rectangular below, rhomboid‑hexagonal to elongate rhomboid above. Setae elongate, straight or flexuose. Capsules subcylindrical, clavate, pyriform, subglobose or ovoid, erect or nodding to pendent, symmetric or curved, smooth, neck well-developed; opercula conic‑apiculate; annuli large, compound, deciduous; stomata superficial; peristome diplolepidous, exostome teeth 16, narrowly triangular, papillose, yellow to brown; endostome yellowish hyaline, lightly papillose, basal membrane well‑developed, segments keeled, narrowly or broadly perforate, cilia 2‑3, nodose or appendiculate, rudimentary or absent. Spores spherical, smooth or papillose. Calyptrae cucullate, smooth, naked.



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